LUQ LTER DATA SETS DOCUMENTATION FORM

ON-LINE VERSION

A DATA SET is a series of observations collected by the same methodology. Each data set should have documentation sufficient for someone unfamiliar with the research to replicate the study. Data sets may be broken into subsets (data files) that are discrete in space and time, in that order. The documentation for a data set should include all spatial and temporal subdivisions of the data.

(Data, Abstract, Methods, Variables)

NOTES:

PERSON(S) COMPLETING THIS FORM: E-MAIL ADDRESS:

Christopher P. Bloch

cbloch@bridgew.edu


DATA SET IDENTIFIER: El Verde Grid invertebrate data

PROJECT TITLE: Community Ecology of Land Snails Survey (Long-term population dynamics of snails in the tabonuco forest)

PROJECT DESCRIPTION: Terrestrial gastropods and walking sticks were sampled at 40 points on the Luquillo Forest Dynamics Plot at El Verde twice annually from 1991-2002 to investigate long-term population and community dynamics and response to disturbance.

Densities

Densities of invertebrates and apparency of plant species are estimated to investigate the effects of disturbance on long-term population dynamics, habitat associations, and spatial distributions.

Habitat

To investigate habitat associations of terrestrial invertebrates, apparency of 17 plant groups (species, genera, families) from 0-3 m from the ground was estimated at each of the sampling localities where invertebrates were surveyed.

LTER CORE AREAS:(Annotate all that apply)
Population Dynamics

Disturbance Patterns

LEF LTER 1 RESEARCH TOPIC: (Annotate all that apply)
Recovery After Disturbance

Spatial and Temporal Distribution of Populations

We define a data file as a component of a data set. A data set can have only one data file or more. Basically, different data files have different data structures or format .
DATA SET FILES (SUBSETS):

Data File No.

Data File Identifier

On-Line Filename

Starting Date

Periodicity of sample

End Period

1 Big Grid Snail MNKA
(Snail Abundance (MNKA))
BigGridSnailCaptures1991-2007.txt June, 1992 Seasonal (Wet, Dry)

June 29, 2007

2

Walking Stick MNKA
(Walking Stick Abundance Data)

WalkingStickMNKA1992-2004.txt June, 1991 Seasonal (Wet, Dry) June, 2004
3

BG Snail M-R95W
(Mark-Recapture Data from Wet Season 1995)

BGSnailM-R95W132-229RECAP.txt June, 1995 Wet season June, 1995
3a BG Snail M-R95W
(Mark-Recapture Data from Wet Season 1995)
BGSnailM-R95W231-328RECAP.txt June, 1995 Wet season June, 1995
3b BG Snail M-R95W
(Mark-Recapture Data from Wet Season 1995)
BGSnailM-R95W330-422RECAP.txt June, 1995 Wet season June, 1995
3c BG Snail M-R95W
(Mark-Recapture Data from Wet Season 1995)
BGSnailM-R95W37-130RECAP.txt June, 1995 Wet season June, 1995
4

BG Snail M-R96D
(Mark-Recapture Data from Dry Season 1996)

BGSnailM-R96D132-229RECAP.txt March, 1996 Dry season March, 1996
4a BG Snail M-R96D
(Mark-Recapture Data from Dry Season 1996)
BGSnailM-R96D231-328RECAP.txt March, 1996 Dry season March, 1996
4b BG Snail M-R96D
(Mark-Recapture Data from Dry Season 1996)
BGSnailM-R96D330-422RECAP.txt March, 1996 Dry season March, 1996
4c BG Snail M-R96D
(Mark-Recapture Data from Dry Season 1996)
BGSnailM-R96D37-130RECAP.txt March, 1996 Dry season March, 1996
5

BG Snail M-R96W
(Mark-Recapture Data from Wet Season 1996)

BGSnailM-R96W132-229RECAP.txt June, 1996 Wet season June, 1996
5a BG Snail M-R96W
(Mark-Recapture Data from Wet Season 1996)
BGSnailM-R96W231-328RECAP.txt June, 1996 Wet season June, 1996
5b BG Snail M-R96W
(Mark-Recapture Data from Wet Season 1996)
BGSnailM-R96W330-422RECAP.txt June, 1996 Wet season June, 1996
5c BG Snail M-R96W
(Mark-Recapture Data from Wet Season 1996)
BGSnailM-R96W37-130RECAP.txt June, 1996 Wet season June, 1996
6

BG Snail M-R97D
(Mark-Recapture Data from Dry Season 1997)

BGSnailM-R97D132-229RECAP.txt March, 1997 Dry season March, 1997
6a BG Snail M-R97D
(Mark-Recapture Data from Dry Season 1997)
BGSnailM-R97D231-328RECAP.txt March, 1997 Dry season March, 1997
6b BG Snail M-R97D
(Mark-Recapture Data from Dry Season 1997)
BGSnailM-R97D330-422RECAP.txt March, 1997 Dry season March, 1997
6c BG Snail M-R97D
(Mark-Recapture Data from Dry Season 1997)
BGSnailM-R97D37-130RECAP.txt March, 1997 Dry season March, 1997
7

BG Snail M-R97W
(Mark-Recapture Data from Wet Season 1997)

BGSnailM-R97W132-229RECAP.txt June, 1997 Wet season June, 1997
7a BG Snail M-R97W
(Mark-Recapture Data from Wet Season 1997)
BGSnailM-R97W231-328RECAP.txt June, 1997 Wet season June, 1997
7b BG Snail M-R97W
(Mark-Recapture Data from Wet Season 1997)
BGSnailM-R97W330-422RECAP.txt June, 1997 Wet season June, 1997
7c BG Snail M-R97W
(Mark-Recapture Data from Wet Season 1997)
BGSnailM-R97W37-130RECAP.txt June, 1997 Wet season June, 1997
8

BG Snail M-R98D
(Mark-Recapture Data from Dry Season 1998)

BGSnailM-R98D132-229RECAP.txt March, 1998 Dry season March, 1998
8a BG Snail M-R98D
(Mark-Recapture Data from Dry Season 1998)
BGSnailM-R98D231-328RECAP.txt March, 1998 Dry season March, 1998
8b BG Snail M-R98D
(Mark-Recapture Data from Dry Season 1998)
BGSnailM-R98D330-422RECAP.txt March, 1998 Dry season March, 1998
8c BG Snail M-R98D
(Mark-Recapture Data from Dry Season 1998)
BGSnailM-R98D37-130RECAP.txt March, 1998 Dry season March, 1998
9

BG Snail M-R98W
(Mark-Recapture Data from Wet Season 1998)

BGSnailM-R98W132-229RECAP.txt June, 1998 Wet season June, 1998
9a BG Snail M-R98W
(Mark-Recapture Data from Wet Season 1998)
BGSnailM-R98W231-328RECAP.txt June, 1998 Wet season June, 1998
9b BG Snail M-R98W
(Mark-Recapture Data from Wet Season 1998)
BGSnailM-R98W330-422RECAP.txt June, 1998 Wet season June, 1998
9c BG Snail M-R98W
(Mark-Recapture Data from Wet Season 1998)
BGSnailM-R98W37-130RECAP.txt June, 1998 Wet season June, 1998
10

BG Snail M-R99D
(Mark-Recapture Data from Dry Season 1999)

BGSnailM-R99D132-229RECAP.txt March, 1999 Dry season March, 1999
10a

BG Snail M-R99D
(Mark-Recapture Data from Dry Season 1999)

BGSnailM-R99D231-328RECAP.txt March, 1999 Dry season March, 1999
10b BG Snail M-R99D
(Mark-Recapture Data from Dry Season 1999)
BGSnailM-R99D330-422RECAP.txt March, 1999 Dry season March, 1999
10c BG Snail M-R99D
(Mark-Recapture Data from Dry Season 1999)
BGSnailM-R99D37-130RECAP.txt March, 1999 Dry season March, 1999
11 BG Snail M-R99W
(Mark-Recapture Data from Wet Season 1999)
BGSnailM-R99W132-229RECAP.txt June, 1998 Wet season June, 1998
11a BG Snail M-R99W
(Mark-Recapture Data from Wet Season 1999)
BGSnailM-R99W231-328RECAP.txt June, 1998 Wet season June, 1998
11b BG Snail M-R99W
(Mark-Recapture Data from Wet Season 1999)
BGSnailM-R99W330-422RECAP.txt June, 1998 Wet season June, 1998
11c BG Snail M-R99W
(Mark-Recapture Data from Wet Season 1999)
BGSnailM-R99W37-130RECAP.txt June, 1998 Wet season June, 1998
12

BG Snail M-R00D
(Mark-Recapture Data from Dry Season 2000)

BGSnailM-R00D132-229RECAP.txt March, 2000 Dry season March, 2000
12a BG Snail M-R00D
(Mark-Recapture Data from Dry Season 2000)
BGSnailM-R00D231-328RECAP.txt March, 2000 Dry season March, 2000
12b BG Snail M-R00D
(Mark-Recapture Data from Dry Season 2000)
BGSnailM-R00D330-422RECAP.txt March, 2000 Dry season March, 2000
12c BG Snail M-R00D
(Mark-Recapture Data from Dry Season 2000)
BGSnailM-R00D37-130RECAP.txt March, 2000 Dry season March, 2000
13

BG Snail M-R00W
(Mark-Recapture Data from Wet Season 2000)

BGSnailM-R00W132-229RECAP.txt July, 2000 Wet season July, 2000
13a BG Snail M-R00W
(Mark-Recapture Data from Wet Season 2000)
BGSnailM-R00W231-328RECAP.txt July, 2000 Wet season July, 2000
13b BG Snail M-R00W
(Mark-Recapture Data from Wet Season 2000)
BGSnailM-R00W330-422RECAP.txt July, 2000 Wet season July, 2000
13c BG Snail M-R00W
(Mark-Recapture Data from Wet Season 2000)
BGSnailM-R00W37-130RECAP.txt July, 2000 Wet season July, 2000
14 BG Snail M-R01D
(Mark-Recapture Data from Dry Season 2001)
BGSnailM-R01D37-130RECAP.txt March, 2001 Dry season March, 2001
14a BG Snail M-R01D
(Mark-Recapture Data from Dry Season 2001)
BGSnailM-R01D132-229RECAP.txt March, 2001 Dry season March, 2001
14b BG Snail M-R01D
(Mark-Recapture Data from Dry Season 2001)
BGSnailM-R01D231-328RECAP.txt March, 2001 Dry season March, 2001
14c BG Snail M-R01D
(Mark-Recapture Data from Dry Season 2001)
BGSnailM-R01D330-422RECAP.txt March, 2001 Dry season March, 2001
15 BG Snail M-R01D
(Mark-Recapture Data from Wet Season 2001)
BGSnailM-R01W37-130RECAP.txt July, 2001 Wet season July, 2001
15a BG Snail M-R01D
(Mark-Recapture Data from Wet Season 2001)
BGSnailM-R01W132-229RECAP.txt July, 2001 Wet season July, 2001
15b BG Snail M-R01D
(Mark-Recapture Data from Wet Season 2001)
BGSnailM-R01W231-328RECAP.txt July, 2001 Wet season July, 2001
15c BG Snail M-R01D
(Mark-Recapture Data from Wet Season 2001)
BGSnailM-R01W330-422RECAP.txt July, 2001 Wet season July, 2001
16 BG Snail M-R02D
(Mark-Recapture Data from Dry Season 2002)
BGSnailM-R02D37-130RECAP.txt May, 2002 Dry Season May, 2002
16a BG Snail M-R02D
(Mark-Recapture Data from Dry Season 2002)
BGSnailM-R02D132-229RECAP.txt May, 2002 Dry Season May, 2002
16b BG Snail M-R02D
(Mark-Recapture Data from Dry Season 2002)
BGSnailM-R02D231-328RECAP.txt May, 2002 Dry Season May, 2002
16c BG Snail M-R02D
(Mark-Recapture Data from Dry Season 2002)
BGSnailM-R02D330-422RECAP.txt May, 2002 Dry Season May, 2002
17 BG Snail M-R01D
(Mark-Recapture Data from Wet Season 2002)
BGSnailM-R02W37-130RECAP.txt July, 2002 Wet season July, 2002
17a BG Snail M-R01D
(Mark-Recapture Data from Wet Season 2002)
BGSnailM-R02W132-229RECAP.txt July, 2002 Wet season July, 2002
17b BG Snail M-R01D
(Mark-Recapture Data from Wet Season 2002)
BGSnailM-R02W231-328RECAP.txt July, 2002 Wet season July, 2002
17c BG Snail M-R01D
(Mark-Recapture Data from Wet Season 2002)
BGSnailM-R02W330-422RECAP.txt July, 2002 Wet season July, 2002
18

BG Snail M-R95W
(Mark-Recapture Totals from Wet Season 1995)

BGSnailM-R95W132-229TOTALS.txt June, 1995 Wet season June, 1995
18a BG Snail M-R95W
(Mark-Recapture Totals from Wet Season 1995)
BGSnailM-R95W231-1928TOTALS.txt June, 1995 Wet season June, 1995
18b BG Snail M-R95W
(Mark-Recapture Totals from Wet Season 1995)
BGSnailM-R95W330-422TOTALS.txt June, 1995 Wet season June, 1995
18c BG Snail M-R95W
(Mark-Recapture Totals from Wet Season 1995)
BGSnailM-R95W37-130TOTALS.txt June, 1995 Wet season June, 1995
19

BG Snail M-R96D
(Mark-Recapture Totals from Dry Season 1996)

BGSnailM-R96D132-229TOTALS.txt March, 1996 Dry season March, 1996
19a BG Snail M-R96D
(Mark-Recapture Totals from Dry Season 1996)
BGSnailM-R96D231-328TOTALS.txt March, 1996 Dry season March, 1996
19b BG Snail M-R96D
(Mark-Recapture Totals from Dry Season 1996)
BGSnailM-R96D330-422TOTALS.txt March, 1996 Dry season March, 1996
19c BG Snail M-R96D
(Mark-Recapture Totals from Dry Season 1996)
BGSnailM-R96D37-130TOTALS.txt March, 1996 Dry season March, 1996
20

BG Snail M-R96W
(Mark-Recapture Totals from Wet Season 1996)

BGSnailM-R96W132-229TOTALS.txt June, 1996 Wet season June, 1996
20a BG Snail M-R96W
(Mark-Recapture Totals from Wet Season 1996)
BGSnailM-R96W231-328TOTALS.txt June, 1996 Wet season June, 1996
20b BG Snail M-R96W
(Mark-Recapture Totals from Wet Season 1996)
BGSnailM-R96W330-422TOTALS.txt June, 1996 Wet season June, 1996
20c BG Snail M-R96W
(Mark-Recapture Totals from Wet Season 1996)
BGSnailM-R96W37-130TOTALS.txt June, 1996 Wet season June, 1996
21

BG Snail M-R97D
(Mark-Recapture Totals from Dry Season 1997)

BGSnailM-R97D132-229TOTALS.txt March, 1997 Dry season March, 1997
21a BG Snail M-R97D
(Mark-Recapture Totals from Dry Season 1997)
BGSnailM-R97D231-328TOTALS.txt March, 1997 Dry season March, 1997
21b BG Snail M-R97D
(Mark-Recapture Totals from Dry Season 1997)
BGSnailM-R97D330-422TOTALS.txt March, 1997 Dry season March, 1997
21c BG Snail M-R97D
(Mark-Recapture Totals from Dry Season 1997)
BGSnailM-R97D37-130TOTALS.txt March, 1997 Dry season March, 1997
22

BG Snail M-R97W
(Mark-Recapture Totals from Wet Season 1997)

BGSnailM-R97W132-229TOTALS.txt June, 1997 Wet season June, 1997
22a BG Snail M-R97W
(Mark-Recapture Totals from Wet Season 1997)
BGSnailM-R97W231-328TOTALS.txt June, 1997 Wet season June, 1997
22b BG Snail M-R97W
(Mark-Recapture Totals from Wet Season 1997)
BGSnailM-R97W330-422TOTALS.txt June, 1997 Wet season June, 1997
22c BG Snail M-R97W
(Mark-Recapture Totals from Wet Season 1997)
BGSnailM-R97W37-130TOTALS.txt June, 1997 Wet season June, 1997
23

BG Snail M-R98D
(Mark-Recapture Totals from Dry Season 1998)

BGSnailM-R98D132-229TOTALS.txt March, 1998 Dry season March, 1998
23a BG Snail M-R98D
(Mark-Recapture Totals from Dry Season 1998)
BGSnailM-R98D231-328TOTALS.txt March, 1998 Dry season March, 1998
23b BG Snail M-R98D
(Mark-Recapture Totals from Dry Season 1998)
BGSnailM-R98D330-422TOTALS.txt March, 1998 Dry season March, 1998
23c BG Snail M-R98D
(Mark-Recapture Totals from Dry Season 1998)
BGSnailM-R98D37-130TOTALS.txt March, 1998 Dry season March, 1998
24

BG Snail M-R98W
(Mark-Recapture Totals from Wet Season 1998)

BGSnailM-R98W132-229TOTALS.txt June, 1998 Wet season June, 1998
24a BG Snail M-R98W
(Mark-Recapture Totals from Wet Season 1998)
BGSnailM-R98W231-328TOTALS.txt June, 1998 Wet season June, 1998
24b BG Snail M-R98W
(Mark-Recapture Totals from Wet Season 1998)
BGSnailM-R98W330-422TOTALS.txt June, 1998 Wet season June, 1998
24c BG Snail M-R98W
(Mark-Recapture Totals from Wet Season 1998)
BGSnailM-R98W37-130TOTALS.txt June, 1998 Wet season June, 1998
25

BG Snail M-R99D
(Mark-Recapture Totals from Dry Season 1999)

BGSnailM-R99D132-229TOTALS.txt March, 1999 Dry season March, 1999
25a

BG Snail M-R99D
(Mark-Recapture Totals from Dry Season 1999)

BGSnailM-R99D231-328TOTALS.txt March, 1999 Dry season March, 1999
25b BG Snail M-R99D
(Mark-Recapture Totals from Dry Season 1999)
BGSnailM-R99D330-422TOTALS.txt March, 1999 Dry season March, 1999
25c BG Snail M-R99D
(Mark-Recapture Totals from Dry Season 1999)
BGSnailM-R99D37-130TOTALS.txt March, 1999 Dry season March, 1999
26 BG Snail M-R99W
(Mark-Recapture Totals from Wet Season 1999)
BGSnailM-R99W132-229TOTALS.txt June, 1998 Wet season June, 1998
26a BG Snail M-R99W
(Mark-Recapture Totals from Wet Season 1999)
BGSnailM-R99W231-328TOTALS.txt June, 1998 Wet season June, 1998
26b BG Snail M-R99W
(Mark-Recapture Totals from Wet Season 1999)
BGSnailM-R99W330-422TOTALS.txt June, 1998 Wet season June, 1998
26c BG Snail M-R99W
(Mark-Recapture Totals from Wet Season 1999)
BGSnailM-R99W37-130TOTALS.txt June, 1998 Wet season June, 1998
27

BG Snail M-R00D
(Mark-Recapture Totals from Dry Season 2000)

BGSnailM-R00D132-229TOTALS.txt March, 2000 Dry season March, 2000
27a BG Snail M-R00D
(Mark-Recapture Totals from Dry Season 2000)
BGSnailM-R00D231-328TOTALS.txt March, 2000 Dry season March, 2000
27b BG Snail M-R00D
(Mark-Recapture Totals from Dry Season 2000)
BGSnailM-R00D330-422TOTALS.txt March, 2000 Dry season March, 2000
27c BG Snail M-R00D
(Mark-Recapture Totals from Dry Season 2000)
BGSnailM-R00D37-130TOTALS.txt March, 2000 Dry season March, 2000
28

BG Snail M-R00W
(Mark-Recapture Totals from Wet Season 2000)

BGSnailM-R00W132-229TOTALS.txt July, 2000 Wet season July, 2000
28a BG Snail M-R00W
(Mark-Recapture Totals from Wet Season 2000)
BGSnailM-R00W231-328TOTALS.txt July, 2000 Wet season July, 2000
28b BG Snail M-R00W
(Mark-Recapture Totals from Wet Season 2000)
BGSnailM-R00W330-422TOTALS.txt July, 2000 Wet season July, 2000
28c BG Snail M-R00W
(Mark-Recapture Totals from Wet Season 2000)
BGSnailM-R00W37-130TOTALS.txt July, 2000 Wet season July, 2000
29 BG Snail M-R01D
(Mark-Recapture Totals from Dry Season 2001)
BGSnailM-R01D37-130TOTALS.txt March, 2001 Dry season March, 2001
29a BG Snail M-R01D
(Mark-Recapture Totals from Dry Season 2001)
BGSnailM-R01D132-229TOTALS.txt March, 2001 Dry season March, 2001
29b BG Snail M-R01D
(Mark-Recapture Totals from Dry Season 2001)
BGSnailM-R01D231-328TOTALS.txt March, 2001 Dry season March, 2001
29c BG Snail M-R01D
(Mark-Recapture Totals from Dry Season 2001)
BGSnailM-R01D330-422TOTALS.txt March, 2001 Dry season March, 2001
30 BG Snail M-R01D
(Mark-Recapture Totals from Wet Season 2001)
BGSnailM-R01W37-130TOTALS.txt July, 2001 Wet season July, 2001
30a BG Snail M-R01D
(Mark-Recapture Totals from Wet Season 2001)
BGSnailM-R01W132-229TOTALS.txt July, 2001 Wet season July, 2001
30b BG Snail M-R01D
(Mark-Recapture Totals from Wet Season 2001)
BGSnailM-R01W231-328TOTALS.txt July, 2001 Wet season July, 2001
30c BG Snail M-R01D
(Mark-Recapture Totals from Wet Season 2001)
BGSnailM-R01W330-422TOTALS.txt July, 2001 Wet season July, 2001
31 BG Snail M-R02D
(Mark-Recapture Totals from Dry Season 2002)
BGSnailM-R02D37-130TOTALS.txt May, 2002 Dry Season May, 2002
31a BG Snail M-R02D
(Mark-Recapture Totals from Dry Season 2002)
BGSnailM-R02D132-229TOTALS.txt May, 2002 Dry Season May, 2002
31b BG Snail M-R02D
(Mark-Recapture Totals from Dry Season 2002)
BGSnailM-R02D231-328TOTALS.txt May, 2002 Dry Season May, 2002
31c BG Snail M-R02D
(Mark-Recapture Totals from Dry Season 2002)
BGSnailM-R02D330-422TOTALS.txt May, 2002 Dry Season May, 2002
32 BG Snail M-R02D
(Mark-Recapture Totals from Wet Season 2002)
BGSnailM-R02W37-130TOTALS.txt July, 2002 Wet season July, 2002
32a BG Snail M-R02D
(Mark-Recapture Totals from Wet Season 2002)
BGSnailM-R02W132-229TOTALS.txt July, 2002 Wet season July, 2002
32b BG Snail M-R02D
(Mark-Recapture Totals from Wet Season 2002)
BGSnailM-R02W231-328TOTALS.txt July, 2002 Wet season July, 2002
32c BG Snail M-R02D
(Mark-Recapture Totals from Wet Season 2002)
BGSnailM-R02W330-422TOTALS.txt July, 2002 Wet season July, 2002
33 BG Snail M-R03D
(Mark-Recapture Data from Dry Season 2003)
BGSnailM-R03D37-130RECAP.txt May, 2003 Dry Season May, 2003
33a BG Snail M-R03D
(Mark-Recapture Data from Dry Season 2003)
BGSnailM-R03D132-229RECAP.txt May, 2003 Dry Season May, 2003
33b BG Snail M-R03D
(Mark-Recapture Data from Dry Season 2003)
BGSnailM-R03D231-328RECAP.txt May, 2003 Dry Season May, 2003
33c BG Snail M-R03D
(Mark-Recapture Data from Dry Season 2003)
BGSnailM-R03D330-422RECAP.txt May, 2003 Dry Season May, 2003
34 BG Snail M-R03D
(Mark-Recapture Data from Wet Season 2003)
BGSnailM-R03W37-130RECAP.txt July, 2003 Wet season July, 2003
34a BG Snail M-R03D
(Mark-Recapture Data from Wet Season 2003)
BGSnailM-R03W132-229RECAP.txt July, 2003 Wet season July, 2003
34b BG Snail M-R03D
(Mark-Recapture Data from Wet Season 2003)
BGSnailM-R03W231-328RECAP.txt July, 2003 Wet season July, 2003
34c BG Snail M-R03D
(Mark-Recapture Data from Wet Season 2003)
BGSnailM-R03W330-422RECAP.txt July, 2003 Wet season July, 2003
35 BG Snail M-R03D
(Mark-Recapture Totals from Dry Season 2003)
BGSnailM-R03D37-130TOTALS.txt May, 2003 Dry Season May, 2003
35a BG Snail M-R03D
(Mark-Recapture Totals from Dry Season 2003)
BGSnailM-R03D132-229TOTALS.txt May, 2003 Dry Season May, 2003
35b BG Snail M-R03D
(Mark-Recapture Totals from Dry Season 2003)
BGSnailM-R03D231-328TOTALS.txt May, 2003 Dry Season May, 2003
35c BG Snail M-R03D
(Mark-Recapture Totals from Dry Season 2003)
BGSnailM-R03D330-422TOTALS.txt May, 2003 Dry Season May, 2003
36 BG Snail M-R03D
(Mark-Recapture Totals from Wet Season 2003)
BGSnailM-R03W37-130TOTALS.txt July, 2003 Wet season July, 2003
36a BG Snail M-R03D
(Mark-Recapture Totals from Wet Season 2003)
BGSnailM-R03W132-229TOTALS.txt July, 2003 Wet season July, 2003
36b BG Snail M-R03D
(Mark-Recapture Totals from Wet Season 2003)
BGSnailM-R03W231-328TOTALS.txt July, 2003 Wet season July, 2003
36c BG Snail M-R03D
(Mark-Recapture Totals from Wet Season 2003)
BGSnailM-R03W330-422TOTALS.txt July, 2003 Wet season July, 2003

RESEARCH LOCATION: Luquillo Forest Dynamics Plots, El Verde

INVESTIGATORS:

PRINCIPAL INVESTIGATORS E-MAIL address

Christopher P. Bloch

cbloch@bridgew.edu

OTHER RESEARCHERS E-MAIL address

Michael Willig

mwillig@lternet.edu

Javier Alvarez javier_alvariz@mail.fws.gov
John Cary  
Stephen Cox stephen.cox@ttu.edu
Michelle Wolf mwolf@montana.edu
Donald Yee dayee@ilstu.edu

CONTACT PERSONS E-MAIL address Phone Number (Include area code)

C. Bloch

cbloch@bridgew.edu

(508)531-2087

SOURCE OF FUNDING (SPONSOR): NSF (Luquillo LTER), U.S. Dept. of Energy (Oak Ridge Associated Universities)

DATA SET ABSTRACT: The data set consists of 13 files: (1) abundance data for the walking stick, Lamponius portoricensis, (2) estimates of abundance (Minimum Number Known Alive) for 17 species of terrestrial snails, and (3) 16 files containing data on size and mark-recapture estimates of population size for the snails Caracolus caracolla and Nenia tridens from Wet Season 1995 to Wet Season 2000. Note: MNKA estimates are not always identical to the number of individuals indicated in the Mark-Recapture data, because some individuals (e.g., those that were lost before being marked) had to be excluded from calculations for Mark-Recapture estimates of abundance. Each of the 16 files contain the data of one of four groups of plots on the Luquillo Forest Dynamics Plot at El Verde: 37-130, 132-229, 231-328, or 330-422, which is identified in the file name along with the years the samples were taken.

In October 2004, the 16 files containing data on size and mark-recapture estimates of population size for the snails Caracolus caracolla and Nenia tridens were separated into two files. One contains the recapture data and another contains the total snails sampled and recaptured for each of these two species.

DATA SET METHODS: One hundred sixty points were selected on the Luquillo Forest Dynamics Plot at El Verde. Circular quadrats (r = 3 m) were established at each point. From June 1991 to present, 40 points were sampled four times seasonally for the presence of terrestrial snails and walking sticks, with the following exceptions: (1) in 1995, 1996, and 1997, the wet season surveys comprised 160 points, (2) in 1998, the wet season survey comprised 100 points, (3) walking stick surveys began in the wet season of 1992, (4) walking sticks were sampled only twice per season until 1994, (5) snails were sampled only once in Dry season 1991, twice in Wet season 1991, Wet and Dry seasons 1992, and Wet and Dry seasons of 1993, and three times in Wet and Dry seasons 1994. Sampling at each point consisted of a minimum of two individuals searching all available surfaces within a quadrat (e.g., vegetation, litter, rock, soil) for the presence of snails and walking sticks. Walking sticks were identified to sex whenever possible, with immature or unidentifiable individuals being classified as "juveniles." Exceptions to this classification system are Dry season 1993, Dry season 1996, and Wet season 1996, when no walking sticks were classified to sex. In these cases, all walking sticks are listed in the data file as "juveniles." Beginning in 1995, individuals of C. caracolla and N. tridens were marked with a numbered plastic dot affixed to the shell and measured to the nearest 0.1 mm using dial calipers. All surveys occurred between 19:30 and 03:00 hours to coincide with peak snail activity. Population densities were estimated as Minimum Number Known Alive (MNKA), the maximum number of individuals of each species recorded for a site in each season, and from Wet Season 1995 onward, using the Lincoln-Peterson, Modified Lincoln-Peterson, and Schnabel estimators.

REFERENCES:
Seber, G. A. F. 1982. The estimation of animal abundance and related parameters. Second ed. MacMillan Publishing Co., Inc., New York, New York, USA.

CROSS-REFERENCES (other data sets related to this one): LTERDBAS 41: Habitat selection/Caracolus caracolla and other snails LTERDBAS 42: Grid Habitat Data, Bisley 1994,1999
LTERDBAS 43: Grid Invertebrate Data Bisley 1989-1999

SAMPLE LOCATION:

STORAGE SITES (of data files):

INVESTIGATOR'S ASSIGNED KEYWORDS: snails, walking sticks, population density, disturbance, Hurricane Hugo

LEF LTER OFFICIAL KEYWORDS (See table): LFDP, TABONUCO, DISTURBANCE, HURRICANE, TERRESTRIAL, PEER REVIEW JOURNAL

PUBLICATIONS:

Secrest, M.F. 1995. The impact of Hurricane Hugo on two common tree snails in the Luquillo Experimental Forest of Puerto Rico: a long-term study. M.S. Thesis, Texas Tech University, Lubbock, Texas.

Secrest, M.F., M.R. Willig, and L.L. Peppers. 1996. The legacy of disturbance on habitat associations of terrestrial snails in the Luquillo Experimental Forest, Puerto Rico. Biotropica 28:502-514.

Willig, M.R., M.F. Secrest, J.F. Cary, S.B. Cox, G.R. Camilo, and M.R. Gannon. 1998. Long-term monitoring of snails in the Luquillo Experimental Forest of Puerto Rico: heterogeneity, scale, disturbance, and recovery. In F. Dallmeier and J. Comisky, editors. Forest Biodiversity in North, Central, and South America and the Caribbean: Research and Monitoring. UNESCO and The Parthenon Press, Carnforth, Lancashire, UK. Pp. 293-322.

Bloch, C.P. 2004. Long-term responses of snail populations and communities to disturbance in the Luquillo Experimental Forest, Puerto Rico. Ph.D. Dissertation. Texas Tech University, Lubbock, TX.

Willig, M.R., C.P. Bloch, N. Brokaw, C. Higgins, J. Thompson, and C. Zimmermann. 2007. Cross-scale responses of biodiversity to hurricane and anthropogenic disturbance in a tropical forest. Ecosystems 10: 834-838.

Bloch, C.P., C.L. Higgins, and M.R. Willig. 2007. Effects of large-scale disturbance on community structure:
temporal trends in nestedness. Oikos 116: 396-406.

Bloch, C.P., and M.R. Willig. 2006. Context-dependence of long-term responses of terrestrial gastropod
populations to large-scale disturbance. Journal of Tropical Ecology 22: 111-122.

DISSEMINATION: UNRESTRICTED

REASONS TO RESTRICT DATA IN THIS DATA SET BEYOND ITS TWO YEAR POLICY PERIOD*:

*WILL HAVE TO BE APPROVED BY LTER PRINCIPAL INVESTIGATORS: J. ZIMMERMAN, A. LUGO , D.J. LODGE

SITES DESCRIPTIONS:

Geographical positional system (GPS) Coordinates for each location:

location

latitude

longitude

     


VARIABLES (ATTRIBUTES):

FILE NAME OR #ABOVE (all in which the variable appears)

1, 3-34 1-34 2 1 1-34 1

ABBREVIATION (as it appears on the data file)

YEAR SEASON YEAR-SEASON RUN POINT DATE

NAME OF VARIABLE

Year Season Year and season Run Point Date

DEFINITION OF VARIABLE

Year of survey (YYYY)

Season when survey was made

Year and season when survey was made (YYS where S = Season with D=Dry and W=Wet ) Identifies which night of sampling the data represent within a
season

Identifies the location on the LFDP (HRP) where sampling was conducted

Date when survey was made (mm/dd/yyyy)

UNIT

N/A N/A N/A N/A N/A N/A

PRECISION

N/A N/A N/A   N/A N/A

RANGE OR LIST OF VALUES

1992... {W= Wet, D=Dry) 92W, 92D, ..., 1 - 4    

DATA TYPE

integer alphanumeric alphanumeric integer integer datetime

MISSING DATA CODES

none none none none none none

VARIABLES (ATTRIBUTES):

FILE NAME OR #ABOVE (all in which the variable appears)

1 1 1 1 1

ABBREVIATION (as it appears on the data file)

AULALT ALCALT ALCSTR CARCAR CARMAR

NAME OF VARIABLE

Population size of captured Austroselenites alticola Population size of captured Alcadia alta Population size of captured Alcadia striata Population size of captured Caracolus caracolla Population size of captured Caracolus marginella

DEFINITION OF VARIABLE

Population size of captured Austroselenites alticola at each point Population size of captured Alcadia alta at each point Population size of captured Alcadia striata at each point Population size of captured Caracolus caracolla at each point Population size of captured Caracolus marginella at each point

UNIT

         

PRECISION

         

RANGE OR LIST OF VALUES

         

DATA TYPE

Integer

Integer Integer

Integer

Integer

MISSING DATA CODES

none none none none none

VARIABLES (ATTRIBUTES):

FILE NAME OR #ABOVE (all in which the variable appears)

1 1 1 1 1 1

ABBREVIATION (as it appears on the data file)

CEPMUS CEPSQU GAENIG LAMGRA MEGCRO NENTRI

NAME OF VARIABLE

Population size of captured Cepolis musicola Population size of captured Cepolis squamosa Population size of captured Gaeotis nigrolineata Population size of captured Lamellaxis gracilis Population size of captured Megalomastoma croceum Population size of captured Nenia tridens

DEFINITION OF VARIABLE

Population size of capturedCepolis musicola at each point Population size of captured Cepolis squamosa at each point Population size of captured Gaeotis nigrolineata at each point Population size of captured Lamellaxis gracilis at each point Population size of captured Megalomastoma croceum at each point Population size of captured Nenia tridens at each point

UNIT

           

PRECISION

           

RANGE OR LIST OF VALUES

           

DATA TYPE

Integer Integer Integer Integer Integer Integer

MISSING DATA CODES

none none none none none none

VARIABLES (ATTRIBUTES):

FILE NAME OR #ABOVE (all in which the variable appears)

1 1 1 1 1

ABBREVIATION (as it appears on the data file)

OLEGRA OLEPLA OBETER POLACU PLAPOR

NAME OF VARIABLE

Population size of captured Oleacina glabra Population size of captured Oleacina playa Population size of captured Obeliscus terebraster Population size of captured Polydontes acutangul Population size of captured Platysuccinea portoricensis

DEFINITION OF VARIABLE

Population size of captured Oleacina glabra at each point Population size of captured Oleacina playa at each point Population size of captured Obeliscus terebraster at each point Population size of captured Polydontes acutangula at each point Population size of captured Platysuccinea portoricensis at each point

UNIT

         

PRECISION

         

RANGE OR LIST OF VALUES

         

DATA TYPE

Integer Integer Integer Integer Integer

MISSING DATA CODES

none none none none none

VARIABLES (ATTRIBUTES):

FILE NAME OR #ABOVE (all in which the variable appears)

1 1 1 1

ABBREVIATION (as it appears on the data file)

SUBOCT VAGOCC UNKNOWN TOTABU

NAME OF VARIABLE

Population size of captured Subulina octana Population size of captured Vaginulus occidentalisf Population size of snails of UNKNOWN species Total abundance (MNKA)

DEFINITION OF VARIABLE

Population size of captured Subulina octana at each point Population size of captured Vaginulus occidentalis at each point Population size of captured snails of unknown species at each point Total abundance (MNKA) of all snails

UNIT

       

PRECISION

       

RANGE OR LIST OF VALUES

       

DATA TYPE

Integer Integer Integer Integer

MISSING DATA CODES

none none none none

VARIABLES (ATTRIBUTES):

FILE NAME OR #ABOVE (all in which the variable appears)

2 2 2 2 2

ABBREVIATION (as it appears on the data file)

JUVENILERUN1 JUVENILERUN2 JUVENILERUN3 JUVENILERUN4 MALERUN1

NAME OF VARIABLE

Juvenile Run 1 Juvenile Run 2 Juvenile Run 3 Juvenile Run 4 Male Run 1

DEFINITION OF VARIABLE

Number of juvenile L. portoricensis encountered the first time the point was surveyed. Number of juvenile L. portoricensis encountered the second time the point was surveyed. Number of juvenile L. portoricensis encountered the third time the point was surveyed. Number of juvenile L. portoricensis encountered the fourth time the point was surveyed. Number of adult male L. portoricensis encountered the first time the point was surveyed.

UNIT

N/A N/A N/A N/A N/A

PRECISION

N/A N/A N/A N/A N/A

RANGE OR LIST OF VALUES

         

DATA TYPE

Integer

Integer

Integer

Integer

Integer

MISSING DATA CODES

Blank Blank Blank Blank Blank

VARIABLES (ATTRIBUTES):

FILE NAME OR #ABOVE (all in which the variable appears)

2 2 2 2 2

ABBREVIATION (as it appears on the data file)

MALERUN2 MALERUN3 MALERUN4 FEMALERUN1 FEMALERUN2

NAME OF VARIABLE

Male Run 2 Male Run 3 Male Run 4 Female Run 1 Female Run 2

DEFINITION OF VARIABLE

Number of adult male L. portoricensis encountered the second time the point was surveyed. Number of adult male L. portoricensis encountered the third time the point was surveyed. Number of adult male L. portoricensis encountered the fourth time the point was surveyed. Number of adult female L. portoricensisencountered the first time the point was surveyed. Number of adult female L. portoricensisencountered the second time the point was surveyed.

UNIT

N/A N/A N/A N/A N/A

PRECISION

N/A N/A N/A N/A N/A

RANGE OR LIST OF VALUES

         

DATA TYPE

Integer Integer Integer Integer Integer

MISSING DATA CODES

Blank Blank Blank Blank Blank

VARIABLES (ATTRIBUTES):

FILE NAME OR #ABOVE (all in which the variable appears)

2 2 2 2 2

ABBREVIATION (as it appears on the data file)

FEMALERUN3 FEMALERUN4 TOTALJUVENILE TOTALMALE TOTALFEMALE

NAME OF VARIABLE

Female Run 3 Female Run 4 Total Juvenile Total Male Total Female

DEFINITION OF VARIABLE

Number of adult female L. portoricensis encountered the third time the point was surveyed. Number of adult female L. portoricensis encountered the fourth time the point was surveyed. Total number of adult juvenile L. portoricensis encountered throughout the season. Total number of adult male L. portoricensis encountered throughout the season. Total number of adult female L. portoricensis encountered throughout the season.

UNIT

N/A N/A N/A N/A N/A

PRECISION

N/A N/A N/A N/A N/A

RANGE OR LIST OF VALUES

         

DATA TYPE

Integer Integer Integer Integer Integer

MISSING DATA CODES

Blank Blank none none none

VARIABLES (ATTRIBUTES):

FILE NAME OR #ABOVE (all in which the variable appears)

2 2 2 2 2

ABBREVIATION (as it appears on the data file)

TOTALADULT TOTALALL MNKAJUVENILE MNKAADULT MNKATOTAL

NAME OF VARIABLE

Total Adult Total Total Minimum Number of Juvenile known alive Minimum Number of Adult known alive Minimum Number of Total known alive

DEFINITION OF VARIABLE

Sum of Total Male and Total Female L. portoricensis encountered throughout the season. Total number of L. portoricensis encountered throughout the season.

Maximum number of juvenile L. portoricensis individuals found at a particular point during the season of interest.

Maximum number of adult L. portoricensisindividuals found at a particular point during the season of interest. Minimum Number of total L. portoricensisindividuals found at a particular point during the season of interest.

UNIT

N/A N/A N/A N/A N/A

PRECISION

N/A N/A N/A N/A N/A

RANGE OR LIST OF VALUES

         

DATA TYPE

Integer

Integer Integer

Integer

Integer

MISSING DATA CODES

none

none none

none

none

VARIABLES (ATTRIBUTES):

FILE NAME OR #ABOVE (all in which the variable appears)

3-17, 33-34

3-17, 33-34

3-17, 33-34

3-17, 33-34 3-17, 33-34

ABBREVIATION (as it appears on the data file)

RECNUM CCCOLORANDNO CCRUN1_MM, CCRUN2_MM CCRUN3_MM

NAME OF VARIABLE

Assigned record number Color and Number identification of Caracolus caracolla Greatest length of shell of a Caracolus caracolla in survey 1 Greatest length of shell of a Caracolus caracolla in survey 2 Greatest length of shell of a Caracolus caracolla in survey 3

DEFINITION OF VARIABLE

Number assigned to record for data management purposes Number and color of marks placed on shells of a Caracolus caracolla individual for identification  Greatest length of shell of a Caracolus caracolla found in the first sampling period within a season Greatest length of shell of a Caracolus caracolla found in the second sampling period within a season Greatest length of shell of a Caracolus caracolla found in the third sampling period within a season

UNIT

N/A   mm mm mm

PRECISION

         

RANGE OR LIST OF VALUES

1,... R = red, B = blue, W = white, etc.)      

DATA TYPE

Integer Alphanumeric Decimal Decimal Decimal

MISSING DATA CODES

none Empty cells in-dicate that an individual was not captured Empty cells in-dicate that an individual was not captured Empty cells in-dicate that an individual was not captured Empty cells in-dicate that an individual was not captured

VARIABLES (ATTRIBUTES):

FILE NAME OR #ABOVE (all in which the variable appears)

3-17, 33-34

3-17, 33-34

3-17, 33-34

3-17, 33-34 3-17, 33-34

ABBREVIATION (as it appears on the data file)

CCRUN4_MM

CCRECAP2

CCRECAP3

CCRECAP4

NTRECAP2

NAME OF VARIABLE

Greatest length of shell of a Caracolus caracolla in survey 4 Recapture event of Caracolus caracolla in survey 2 Recapture event of Caracolus caracolla in survey 3 Recapture event of Caracolus caracolla in survey 4 Recapture event of Nenia tridens in survey 2

DEFINITION OF VARIABLE

Greatest length of shell of a Caracolus caracolla found in the fourth sampling period within a season Whether any individual of Caracolus caracolla captured in survey 2 had been captured previously in the same season Whether any individual of Caracolus caracollacaptured in survey 3 had been captured previously in the same season Whether any individual of Caracolus caracolla captured in survey 4 had been captured previously in the same season Whether any individual of Nenia tridens captured in survey 2 had been captured previously in the same season

UNIT

mm        

PRECISION

         

RANGE OR LIST OF VALUES

 

0 = no recapture,
1 = recapture occured

0 = no recapture,
1 = recapture occured
0 = no recapture,
1 = recapture occured
0 = no recapture,
1 = recapture occured

DATA TYPE

Decimal Integer Integer Integer Integer

MISSING DATA CODES

Empty cells in-dicate that an individual was not captured none none none none

VARIABLES (ATTRIBUTES):

FILE NAME OR #ABOVE (all in which the variable appears)

3-17, 33-34

3-17, 33-34

3-17, 33-34

3-17, 33-34 3-17, 33-34

ABBREVIATION (as it appears on the data file)

NTRECAP3

NTRECAP4RUN

NTCOLORANDNO NTRUN1_MM, NTRUN2_MM,

NAME OF VARIABLE

Recapture event of Nenia tridens in survey 3 Recapture event of Nenia tridens in survey 4 Color and Number identification of Nenia tridens Greatest length of shell of Nenia tridens Greatest length of shell of a Nenia tridens in survey 2

DEFINITION OF VARIABLE

Whether any individual of Nenia tridens captured in survey 3 had been captured previously in the same season Whether any individual of Nenia tridens captured in survey 4 had been captured previously in the same season Number and color of marks placed on shells of a Nenia tridens individual for identification  Greatest length of shell in each sampling period within a season Greatest length of shell of a Nenia tridensfound in the second sampling period within a season

UNIT

      mm mm

PRECISION

         

RANGE OR LIST OF VALUES

0 = no recapture,
1 = recapture occured
0 = no recapture,
1 = recapture occured
R = red, B = blue, W = white, etc.)    

DATA TYPE

Integer Integer Alphanumeric Decimal Decimal

MISSING DATA CODES

none none Empty cells in-dicate that an individual was not captured Empty cells in-dicate that an individual was not captured Empty cells in-dicate that an individual was not captured

VARIABLES (ATTRIBUTES):

FILE NAME OR #ABOVE (all in which the variable appears)

3-17, 33-34

3-17, 33-34

18-32,35-36

18-32,35-36 18-32,35-36

ABBREVIATION (as it appears on the data file)

NTRUN3_MM, NTRUN4_MM TOTAL SPECIESCODE TOTALTYPE

NAME OF VARIABLE

Greatest length of shell of a Nenia tridens in survey 3 Greatest length of shell of a Nenia tridens in survey 4 Total number of individuals captured Species name code Run or recapture event

DEFINITION OF VARIABLE

Greatest length of shell of a Nenia tridens found in the third sampling period within a season Greatest length of shell of a Nenia tridens found in the fourth sampling period within a season Total number of individuals captured or recapture at a site within a season. For recapture events this field is left blank for the first Run Two letter species code of the species studied Type of activity for which total has been either calculated for a specific point in a sampling period within a season

UNIT

mm        

PRECISION

         

RANGE OR LIST OF VALUES

      NT = Nenia tridens,CC = Caracolus caracolla run = sampling period , recapture = recapture event

DATA TYPE

Decimal   Integer Alphabetic  

MISSING DATA CODES

Empty cells in-dicate that an individual was not captured   none none  

 

COMPUTATIONAL METHODS:  
VARIABLE NAME FORMULA OR DESCRIPTION 
Minimum Number Known Alive Maximum number of individuals found at a particular point during the season of interest.
Lincoln-Peterson Estimate of Population Size N=M/(m/n), where N=population size, n=total individuals captured in a sample, m=recaptured individuals in that sample, and M=total number of marked individuals in the population
Modified Lincoln-Peterson Estimate of Population Size N=M/((m/n)-1), where n=total individuals in a sample +1, m=recaptured individuals in that sample +1, and M=total marked individuals in the population +1
Schnabel Estimate of Population Size N=sum of (M*n) for each time period/sum of m over all time periods, where N,M,m,n are defined as ifor the Lincoln-Peterson estimator.


FOR DATA MANAGER USE ONLY

DATE OF LAST REVIEW: January 24, 2008
DATE OF LAST ENTRY: 2003
STAGE OF DATA SET MANAGEMENT (dates):
RECEIVED SEP 29 2000
ENTERED:
FILED: Oct 4, 2000
ON-LINE:  Oct 4, 2000
REVIEWED BY RESEARCHER
FILING MEDIA:
NAME OF DOCUMENTATION FILE: Lterdb107.Htm; Hrpinverts.Doc
NAME OF ON - LINE CATALOG: LTERDBAS 
RECORD #: 107
DOCUMENT TYPE: magnetic media
PRIORITY TO BE ENTERED: N/A


Rev. date of this form: 8 June 2000