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DATA SET IDENTIFIER: Does the River Continuum Concept apply on a tropical island? Longitudinal variation in a Puerto Rican stream

PROJECT TITLE: Does the River Continuum Concept apply on a tropical island? Longitudinal variation in a Puerto Rican stream (Community Composition of Fresh Water Shrimps)

PROJECT DESCRIPTION: As part of Luquillo LTER efforts to describe how ecological communities of the Luquillo Mountains change with elevation, we sampled stream communities and ecosystem properties at 9 sites along an elevational gradient in the Río Mameyes catchment.

LTER CORE AREAS: Annotate all that apply (See online list)

Population Dynamics

LEF LTER 1 RESEARCH TOPIC: (Annotate all that apply) (See online list)

Recovery-Spat. Dist.

We define a data file as a component of a data set. A data set can have only one data file or more. Basically, different data files have different data structures or format .
DATA SET FILES (SUBSETS):

Data File No.	Data File Identifier	On-Line Filename	Starting Date	Periodicity of sample	End Period
1	Locations of study sites	mameyessites.txt	N/A	N/A	N/A
2	Presence/absence of predatory fishes	predatoryfishes.txt	June 2001	irregular	August 2001
3	Biomass of decapods	decapods.txt	June 2001	irregular	August 2001
4	Biomass of non-decapod invertebrates	nondecapodinverts.txt	June 2001	irregular	August 2001
5	Biomass of macroinvertebrate functional feeding groups	FFGs.txt	June 2001	irregular	August 2001
6	Standing stocks of basal and benthic resources	basalresources.txt	June 2001	irregular	August 2001
7	Proportional representation of habitats	habitatrepresentation.txt	June 2001	irregular	August 2001
8	Density of Sicydium plumieri	gobies.txt	June 2001	irregular	August 2001

RESEARCH LOCATION: Río Mameyes and tributaries (Río de La Mina and an unnamed tributary).

INVESTIGATORS:

PRINCIPAL INVESTIGATORS	E-MAIL ADDRESS:
Effie A. Greathouse	effieg@gmail.com

OTHER RESEARCHERS	E-MAIL ADDRESS:
Catherine M. Pringle	cpringle@uga.edu

CONTACT PERSONS	Phone Number (Include area code)	E-MAIL ADDRESS:
Effie A. Greathouse	916-332-5771 (Whitney Masonry)	effieg@gmail.com

SOURCE OF FUNDING (SPONSOR): National Science Foundation (NSF) Luquillo LTER (BSR 8811902, DEB9411973, DEB0080538, DEB0218039), the NSF Doctoral Dissertation Improvement Grant program (DEB 0308543), the U.S. Department of Agriculture (grants 10-21-RR551-141, 10-21-RR250-109); and the NSF Graduate Fellowship program (fellowship to EAG)

DATA SET ABSTRACT: We examined whether a tropical stream in Puerto Rico matched predictions of the River Continuum Concept (RCC) for macroinvertebrate functional feeding groups (FFGs). Sampling sites for macroinvertebrates, basal resources, and fishes ranged from headwaters to within 2.5 km of the fourth-order estuary. In a comparison to a model temperate system where RCC predictions generally held, we used catchment area as a measure of stream size in order to examine truncated RCC predictions (i.e., cut off to correspond to the largest stream size sampled in Puerto Rico). Despite dominance of generalist freshwater shrimps, which use more than one feeding mode, RCC predictions held for scrapers, shredders, and predators. Collector-filterers showed a trend opposite that predicted by the RCC, but patterns in basal resources suggest that this is consistent with the central RCC theme: longitudinal distributions of FFGs follow longitudinal patterns in basal resources. Alternatively, the filterer pattern may be explained by fish predation affecting distributions of filter-feeding shrimp. Our results indicate that the RCC generally applies to running waters on tropical islands. However, additional theoretical and field studies across a broad array of stream types should examine whether the RCC needs to be refined to reflect the potential influence of top-down trophic controls on FFG distributions.

DATA SET METHODS: See Greathouse et al. (2006) and Greathouse (2005). (full reference in the "REFERENCES" section below). At each site, we established a reach that was 10 times the channel width and visually identified major habitats. Proportional availability of major habitats was determined using occurrence frequency of mesohabitats (riffles, pools, and runs, based on measured lengths) combined with occurrence frequency of substrates (based on 54 random measurements in each major mesohabitat type). All measurements and samples were taken during base flow from June through August 2001. Epilithic chlorophyll a was sampled using a suction device modified from Loeb (1981). We collected a pooled sample of four individual and randomly located Loeb samples in each of three pools and three riffles at each site. Chlorophyll a samples were brought back to the lab on ice, filtered, frozen, and analyzed fluorometrically according to standard methods (American Public Health Association 1985).

To measure non-shrimp benthic invertebrate densities and standing stocks of very fine (VF, <250 m) and medium fine (MF, 250 m – 1 mm) benthic inorganic matter (BIM) and VF, MF, and coarse (C, >1 mm) benthic organic matter (BOM), three randomly located samples per major habitat were collected and processed using methods modified from Lugthart and Wallace (1992) and Grubaugh et al. (1996). Macrophytes in runs, as well as sand habitats in pools, were sampled using a core of known area. Pools with boulders and interstitial cobble and gravel were sampled using the "benthic block net" method described by Greathouse and Pringle (2005); the benthic block net apparatus functions similarly to a core. Bedrock–boulder cascades were sampled by scrub-brushing invertebrates and benthic matter from a known area into a hand net or into a Surber sampler with foam rubber tied around the frame to create a seal with the substrate. Cobble–coarse gravel riffles were sampled with an unmodified Surber net. In cobble runs, a sample consisted of a single cobble placed into a downstream D net, scrubbed on the bank to remove invertebrates and benthic matter, and measured to determine surface area. Because hand, Surber, and D nets were 250 m mesh, we further modified methods of Lugthart and Wallace (1992) and Grubaugh et al. (1996) by using the Loeb sampler to collect VFBOM and VFBIM in Surber and hand-net samples, and we used the Loeb sampler on nearby cobbles to estimate VFBOM and VFBIM in D-net samples. An additional modification is that we did not separate macrophytes from coarse detritus when measuring standing stocks of CBOM; however, macrophytes only occurred in run and riffle samples at the lowest-elevation site (Hydrilla) and in a single riffle sample at site 3 (filamentous algae). Invertebrate samples were preserved in the field in ethanol.

Samples intended for estimating VFBOM/BIM (from Loeb, core, and benthic block net sampling) were sieved to remove material >250 m before filtration onto pre-ashed, preweighed glass fiber filters (Whatman GF/F, 0.7 m). Nondecapod invertebrate samples were separated into MF and C fractions using nested 1 mm and 250 m sieves before sorting for macroinvertebrates. After sorting, we subsampled the MFBOM/BIM for filtration onto a pre-ashed, pre-weighed glass fiber filter. CBOM and filters of VFBOM/BIM and MFBOM/BIM were dried for 24 h at 50 °C, weighed, ashed for 3 h at 500 °C, and reweighed to determine ash-free dry mass (AFDM) for VFBOM, MFBOM, and CBOM and inorganic dry mass for VFBIM and MFBIM. Non-decapod invertebrates were identified to the lowest practicable level © 2005 NRC Canada 136 Can. J. Fish. Aquat. Sci. Vol. 63, 2006.

We obtained quantitative estimates of shrimp and crab biomass and fish abundance by electroshocking and snorkeling. A minimum of one pool or run habitat and one riffle habitat per site were electroshocked using depletion or removal sampling methods (White et al. 1982; Fièvet et al. 1996, 1999). Block and dip nets were 0.635 cm mesh. Captured shrimps and crabs were identified to genus or species and measured to the nearest millimetre for postorbital carapace length (Hobbs 1991). Electroshocking was ineffective for capturing gobies (Sicydium and Awaous) and sleepers (Eleotridae) in pools, as has been observed by other researchers for Puerto Rico (E. García, US Department of Agriculture Forest Service, Southern Region, Atlanta, GA 30309, USA, and N. Hemphill, Trinity County Resource Conservation District, Weaverville, CA 96093, USA, personal communication). Thus, in one pool or run per site, we snorkeled a known area, counting all gobies and sleepers observed. Visibility was similar across sites. Carapace length– mass regression equations for shrimps and crabs, determined from individuals preserved in ~10% formaldehyde, were used to determine shrimp and crab biomass (Appendix A). In determining biomass of macroinvertebrate functional feeding groups, we apportioned biomass of shrimp taxa equivalently to each known functional feeding group membership.

REFERENCES:
Greathouse, E.A. 2005. Large dams and migratory biota affect tropical stream ecosystems at different scales in Puerto Rico. Ph.D. Dissertation, University of Georgia, Athens.

Greathouse, E. A., and C. M. Pringle. 2006. Does the River Continuum Concept apply on a tropical island? Longitudinal variation in a Puerto Rican stream. Canadian Journal of Fisheries and Aquatic Sciences 63: 134-152

CROSS-REFERENCES (other data sets related to this one):

SAMPLE LOCATION: N/A

STORAGE SITES(of data files): Data is maintained by Effie Greathouse.

INVESTIGATOR'S ASSIGNED KEYWORDS: Decapoda; freshwater shrimps; functional feeding groups; macroinvertebrates

LEF LTER OFFICIAL KEYWORDS (See table): Q MAMEYES, HEADWATER STREAM, RIVER, COMMUNITY COMPOSITION, ALGAE, AQUATIC, DECAPODS, FISH, INSECTS, GASTROPODS, DISSERTATION, PEER REVIEWED JOURNAL

PUBLICATIONS:
Greathouse, E. A., and C. M. Pringle. 2006. Does the River Continuum Concept apply on a tropical island? Longitudinal variation in a Puerto Rican stream. Canadian Journal of Fisheries and Aquatic Sciences 63: 134-152

Greathouse, E. A., and C. M. Pringle. 2005. A sampler for stream macroinvertebrates and organic matter occurring on boulders and bedrock in pools. Ver. Int. Ver. Theor. Angew. Limnol. 29/2: 975-978

RESTRICTED ___ UNRESTRICTED _X_

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)	1, 2, 3, 4, 5, 6, 7, 8	1	1	1
AbbreviationAbbreviation(as it appears on the data file)	Site	Stream/location	latdeg	latmin
NAME OF VARIABLE	Site number	Stream name/location	Latitude degrees	Latitude minutes
DEFINITION OF VARIABLE	A number assigned to each sampling site	The name of the stream site or its location	The degrees of the site's latitude coordinates	The minutes of the site's latitude coordinates
UNIT		n/a	degrees	minutes
PRECISION	{1,.,9}	see data file	{18}	{17,.,22}
DATA TYPE	alphanumeric	other	integer	integer
MISSING DATA CODES	none	none	none	none

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)	1	1	1	1
AbbreviationAbbreviation(as it appears on the data file)	latsec	londeg	lonmin	lonsec
NAME OF VARIABLE	Latitude seconds	Longitude degrees	Longitude minutes	Longitude seconds
DEFINITION OF VARIABLE	The seconds of the site's latitude coordinates	The degrees of the site's longitude coordinates	The minutes of the site's longitude coordinates	The seconds of the site's longitude coordinates
UNIT	seconds	degrees	minutes	seconds
PRECISION	0.01 seconds	absolute	absolute	0.01 seconds
RANGE OR LIST OF VALUES	{5.24,.,55.79}	{65}	{45,.,47}	{0.23,.,59.2}
DATA TYPE	decimal	integer	integer	decimal
MISSING DATA CODES	none	none	none	none

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)	2, 3, 4, 5, 6, 7	2	2	2
AbbreviationAbbreviation(as it appears on the data file)	habitat	angros	agomon	eleotrid
NAME OF VARIABLE	habitat	Anguilla rostrata	Agonostomus monticola	Eleotridae
DEFINITION OF VARIABLE	The habitat that the measurement or observation was taken in; habitats were: riffle (sampled at sites 1-9), pool (sampled at sites 1-8), run (sampled at site 9), cobble (cobble substrate within the run habitat at site 9), hydrilla (Hydrilla macrophytes within the run habitat at site 9)	Presence/absence of Anguilla rostrata in electroshocking samples or observations of the habitat at the site, where x indicates presence and 0 indicates absence	Presence/absence of Agonostomus monticola in electroshocking samples or observations of the habitat at the site, where x indicates presence and 0 indicates absence	Presence/absence of Eleotridae in electroshocking samples or observations of the habitat at the site, where x indicates presence and 0 indicates absence
UNIT	N/A	N/A	N/A	N/A
PRECISION	absolute	absolute	absolute	absolute
RANGE OR LIST OF VALUES	riffle, pool, run, cobble, hydrilla	x, 0	x, 0	x, 0
DATA TYPE	other	logical	logical	logical
MISSING DATA CODES	none	none	none	none

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)	3	3	3	3
AbbreviationAbbreviation(as it appears on the data file)	atyid	xipho	mac	crab
NAME OF VARIABLE	Atyidae biomass	Xiphocaris biomass	Macrobrachium spp. biomass	Crab biomass
DEFINITION OF VARIABLE	Biomass of Atyidae sampled by electroshocking removal methods	Biomass of Xiphocaris elongata sampled by electroshocking removal methods	Biomass of Macrobrachium spp. sampled by electroshocking removal methods	Biomass of crabs (Epilobocera sinuatifrons) sampled by electroshocking removal methods
UNIT	grams ash-free dry mass per square meter	grams ash-free dry mass per square meter	grams ash-free dry mass per square meter	grams ash-free dry mass per square meter
PRECISION	0.1 g AFDM/square m	0.1 g AFDM/square m	0.1 g AFDM/square m	0.1 g AFDM/square m
RANGE OR LIST OF VALUES	{0,.,10.1}	{0,.,1.9}	{0,.,2.9}	{0,.,2.7}
DATA TYPE	decimal	decimal	decimal	decimal
MISSING DATA CODES	none	none	none	none

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)	4	4	4	4
AbbreviationAbbreviation(as it appears on the data file)	phanocerus	otherelmid	blepharicerid	chironomid
NAME OF VARIABLE	Phanocerus spp. biomass	Biomass of non-Phanocerus Elmidae	Blephariceridae biomass	Chironomidae biomass
DEFINITION OF VARIABLE	Biomass of Phanocerus spp. (Elmidae)	Biomass of Elmidae other than Phanocerus spp.	Biomass of Blephariceridae	Biomass of Chironomidae
UNIT	milligrams ash-free dry mass per square meter	milligrams ash-free dry mass per square meter	milligrams ash-free dry mass per square meter	milligrams ash-free dry mass per square meter
PRECISION	1 mg AFDM/square m	1 mg AFDM/square m	1 mg AFDM/square m	1 mg AFDM/square m
RANGE OR LIST OF VALUES	{0,.,6}	{0,.,159}	{0,.,65}	{<1,.,216}
DATA TYPE	integer	integer	integer	integer
MISSING DATA CODES	none	none	none	none

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)	4	4	4	4
AbbreviationAbbreviation(as it appears on the data file)	simuliid	baetid	leptophlebiid	crambid
NAME OF VARIABLE	Simuliidae biomass	Baetidae biomass	Leptophlebiidae biomass	Crambidae biomass
DEFINITION OF VARIABLE	Biomass of Simuliidae	Biomass of Baetidae	Biomass of Leptophlebiidae	Biomass of Crambidae
UNIT	milligrams ash-free dry mass per square meter	milligrams ash-free dry mass per square meter	milligrams ash-free dry mass per square meter	milligrams ash-free dry mass per square meter
PRECISION	1 mg AFDM/square m	1 mg AFDM/square m	1 mg AFDM/square m	1 mg AFDM/square m
RANGE OR LIST OF VALUES	{0,.,31}	{0,.,741}	{0,.,61}	{0,.,124}
DATA TYPE	integer	integer	integer	integer
MISSING DATA CODES	none	none	none	none

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)	4	4	4	5
AbbreviationAbbreviation(as it appears on the data file)	neritina	thiara	other	dscrapers
NAME OF VARIABLE	Biomass of Neritina spp.	Thiara granifera biomass	Biomass of other non-decapod invertebrate taxa	Decapod scraper biomass
DEFINITION OF VARIABLE	Biomass of Neritina spp.	Biomass of Thiara granifera	Biomass of other non-decapod invertebrate taxa (Ceratopogonidae, Corethrellidae, Dixidae, Empididae, Psychodidae, Stratiomyidae, Tabanidae, Caenidae, Corixidae, Coengionidae, Calamoceratidae, Glossosomatidae, Helicopsychidae, Leptoceridae, Polycentropodidae, Psychomyidae, Xiphocentronidae, limpets, Acari, Collembola, Copepoda, Oligochaeta, Ostracoda, Turbellaria)	Decapod biomass assigned to the scraper functional feeding group
UNIT	milligrams ash-free dry mass per square meter	milligrams ash-free dry mass per square meter	milligrams ash-free dry mass per square meter	grams ash-free dry mass per square meter
PRECISION	1 mg AFDM/square m	1 mg AFDM/square m	1 mg AFDM/square m	0.1 g AFDM/square m
RANGE OR LIST OF VALUES	{0,.,299}	{0,.,299}	{<1,.,17}	{<0.1,.,3.3}
DATA TYPE	integer	integer	integer	decimal
MISSING DATA CODES	none	none	none	none

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)	3, 8	3	3	3, 8
AbbreviationAbbreviation(as it appears on the data file)	cancov	rifflevel	poolvel	depth
NAME OF VARIABLE	Canopy cover	Riffle water velocity	Pool water velocity	Water depth
DEFINITION OF VARIABLE	Canopy cover over the stream reach	Mean water velocity in riffles	Mean water velocity in pools	Mean water depth
UNIT	percent	meters per second	meters per second	meters
PRECISION	0.1 percent	0.01 m/s	0.01 m/s	0.01 m
RANGE OR LIST OF VALUES	{0,.,90.5}	{0.29,.,1.02}	{-0.02,.,0.07}	{0.11,.,0.43}
DATA TYPE	decimal	decimal	decimal	decimal
MISSING DATA CODES	none	--- indicates no measurement of water velocity	--- indicates no measurement of water velocity	none

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)	5	5	5	5
AbbreviationAbbreviation(as it appears on the data file)	ngatherers	dfilterers	nfilterers	dpredators
NAME OF VARIABLE	Non-decapod collector-gatherer biomass	Decapod collector-filterer biomass	Non-decapod collector-filterer biomass	Decapod predator biomass
DEFINITION OF VARIABLE	Non-decapod invertebrate biomass assigned to the collector-gatherer functional feeding group	Decapod biomass assigned to the collector-filterer functional feeding group	Non-decapod invertebrate biomass assigned to the collector-filterer functional feeding group	Decapod biomass assigned to the predator functional feeding group
UNIT	milligrams ash-free dry mass per square meter	grams ash-free dry mass per square meter	milligrams ash-free dry mass per square meter	grams ash-free dry mass per square meter
PRECISION	1 mg AFDM/square m	0.1 g AFDM/square m	1 mg AFDM/square m	0.1 g AFDM/square m
RANGE OR LIST OF VALUES	{0.3,.,304}	{0,.,3.9}	{<1,.,299}	{0,.,1.5}
DATA TYPE	integer	decimal	integer	decimal
MISSING DATA CODES	none	none	none	none

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)	5	6	6	6
AbbreviationAbbreviation(as it appears on the data file)	npredators	chl	VFBOM	MFBOM
NAME OF VARIABLE	Non-decapod predator biomass	Chlorophyll a	Very fine benthic organic matter	Medium fine benthic organic matter
DEFINITION OF VARIABLE	Non-decapod invertebrate biomass assigned to the predator functional feeding group	Standing crop of chlorophyll a sampled from rocks (cobble, boulder, bedrock)	Standing stock of very fine (<250 micron) benthic organic matter occurring in non-decapod macroinvertebrate samples	Standing stock of medium (<1mm, >250 micron) benthic organic matter occurring in non-decapod macroinvertebrate samples
UNIT	milligrams ash-free dry mass per square meter	milligrams per square meter	grams ash-free dry mass per square meter	grams ash-free dry mass per square meter
PRECISION	1 mg AFDM/square m	0.1 mg/square m	0.1 g AFDM/square m	0.1 g AFDM/square m
RANGE OR LIST OF VALUES	{<1,.,81}	{0.2,.,15.3}	{0.7,.,73.2}	{<0.1,.,249.0}
DATA TYPE	integer	decimal	decimal	decimal
MISSING DATA CODES	none	--- indicates that chlorophyll a was not sampled from that habitat	none	none

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)	6	6	6	7
AbbreviationAbbreviation(as it appears on the data file)	CBOM	VFBIM	MFBIM	overall
NAME OF VARIABLE	Coarse benthic organic matter	Very fine benthic inorganic matter	Medium fine benthic inorganic matter	proportional habitat representation for decapod sampling
DEFINITION OF VARIABLE	Standing stock of coarse (>1mm) benthic organic matter occurring in non-decapod macroinvertebrate samples	Standing stock of very fine (<250 micron) benthic inorganic matter occurring in non-decapod macroinvertebrate samples	Standing stock of medium (<1mm, >250 micron) benthic inorganic matter occurring in non-decapod macroinvertebrate samples	Proportional representation of sampled mesohabitats (riffle, pool, run); these proportions were used for habitat-weighting of decapod biomass
UNIT	grams ash-free dry mass per square meter	grams ash-free dry mass per square meter	grams ash-free dry mass per square meter	proportion
PRECISION	0.1 g AFDM/square m	0.1 g DM/square m	0.1 g DM/square m	N/A
RANGE OR LIST OF VALUES	{<0.1,.,558.0}	{0.1,.,381.0}	{<0.1,.,6736.0}	{0.22,.,0.78}
DATA TYPE	decimal	decimal	decimal	decimal
MISSING DATA CODES	none	none	none	none

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)	7	7	7	7
AbbreviationAbbreviation(as it appears on the data file)	boulderbedrock	sand	cobble	hydrilla
NAME OF VARIABLE	proportional habitat representation of boulder/bedrock	proportional habitat representation of sand	proportional habitat representation of cobble	proportional habitat representation of Hydrilla macrophytes
DEFINITION OF VARIABLE	Proportional representation of boulder/bedrock substrate at sites where it was sampled for non-decapod invertebrate biomass; these proportions were used for habitat-weighting of non-decapod invertebrate biomass	Proportional representation of sand substrate at sites where it was sampled for non-decapod invertebrate biomass; these proportions were used for habitat-weighting of non-decapod invertebrate biomass	Proportional representation of cobble substrate at sites where it was sampled for non-decapod invertebrate biomass; these proportions were used for habitat-weighting of non-decapod invertebrate biomass	Proportional representation of Hydrilla macrophytes as a substrate at sites where it was sampled for non-decapod invertebrate biomass; these proportions were used for habitat-weighting of non-decapod invertebrate biomass
UNIT	proportion	proportion	proportion	proportion
PRECISION	N/A	N/A	N/A	N/A
RANGE OR LIST OF VALUES	{0.14,.,0.76}	{0.27}	{0.13,.,0.53}	{0.61}
DATA TYPE	decimal	decimal	decimal	decimal
MISSING DATA CODES	--- indicates that boulder/bedrock substrate was not sampled for non-decapod invertebrate biomass at that site/mesohabitat	--- indicates that sand substrate was not sampled for non-decapod invertebrate biomass at that site/mesohabitat	--- indicates that cobble substrate was not sampled for non-decapod invertebrate biomass at that site/mesohabitat	--- indicates that Hydrilla macrophytes were not sampled for non-decapod invertebrate biomass at that site/mesohabitat

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)	8	8
AbbreviationAbbreviation(as it appears on the data file)	poolgoby	rifflegoby
NAME OF VARIABLE	Sicydium density in pools	Sicydium density in riffles
DEFINITION OF VARIABLE	Density of Sicydium plumieri in pool habitat, as estimated by counting S. plumieri while snorkeling a known area	Density of Sicydium plumieri in riffle habitat, as estimated by depletion sampling by electroshocking
UNIT	number of individuals per square meter	numbers of individuals per square meter
PRECISION	0.01 individuals/square m	0.01 individuals/square m
RANGE OR LIST OF VALUES	{0,.,2.93}	{0,.,1.44}
DATA TYPE	decimal	decimal
MISSING DATA CODES	none	none

COMPUTATIONAL METHODS:

Variable Name	Formula

DATE OF LAST REVIEW: June 2, 2006

DATE OF LAST ENTRY: August, 2000

STAGE OF DATA SET MANAGEMENT (dates):
RECEIVED: January 5, 2006
ENTERED: January 28, 2006
FILED ON-LINE: February 4, 2006
REVIEWED BY RESEARCHER:

FILING MEDIA:
NAME OF DOCUMENTATION FILE: CJFASMetadataForm.doc, lterdb142.htm
NAME OF ON - LINE CATALOG: LTERDBAS
RECORD #: 142
DOCUMENT TYPE: binary data only
PRIORITY TO BE ENTERED: N/A

RECEIVED
ENTERED: