LUQ LTER DATA SETS DOCUMENTATION FORM

ON-LINE VERSION

A DATA SET is a series of observations collected by the same methodology. Each data set should have documentation sufficient for someone unfamiliar with the research to replicate the study. Data sets may be broken into subsets (data files) that are discrete in space and time, in that order. The documentation for a data set should include all spatial and temporal subdivisions of the data.

(Data, Abstract, Methods, Variables)

NOTES:

PERSON(S) COMPLETING THIS FORM: E-MAIL ADDRESS:

Larry Woolbright

lwoolbright@lternet.edu

Eda C. Meléndez-Colom

emelendez@lternet.edu

DATA SET IDENTIFIER: Pattern morphology for frogs captured at 9 locations in northeastern Puerto Rico over a 25-year period from 1978 to 2002

PROJECT TITLE: Frog Population Dynamics

PROJECT DESCRIPTION: El Verde Station, located in the tabonuco forest zone on the NW side of the Luquillo Experimental Forest (LEF), has hosted numerous scientific efforts since the 1960s. Thus, a great deal of information exists concerning anuran populations at the site over the past 30 years. Although past surveys did not use consistent techniques, they clearly document the general status of the anuran community of the area. I began working at El Verde in the 1970s and have collected data on the anuran community primarily in the 1980s and 1990s

LTER CORE AREAS: Annotate all that apply (See online list)

Population Dynamics
 

LEF LTER 1 RESEARCH TOPIC: (Annotate all that apply) (See online list)

Recovery-Env. Mon.


We define a data file as a component of a data set. A data set can have only one data file or more. Basically, different data files have different data structures or format .
DATA SET FILES (SUBSETS):

Data File No.

Data File Identifier

On-Line Filename

Starting Date

Periodicity of sample

End Period

1

Sonadora Plot 

SonadoraPlot.txt January 1, 1987 varies August 2, 2002
2 Bisley Stream 2 Plot BisleyStream2Plot.txt June 1, 1988 varies August 2, 2002
3 Bisley Forest Control Plot BisleyForestControlPlot.txt June 1, 1988 varies August 1, 2002
4 Margaret Stewart Plots MargaretStewartPlots.txt January 1, 1984 varies January 31, 1984

RESEARCH LOCATION: Nine locations in northeastern Puerto Rico: 1. LLW’s Sonadora Plot just west of the Sonadora River uphill from the main trail at EVFS, 2. LLW’s Bisley Stream 2 plot in the stream 2 channel, 3. LLW’s Bisley Forest Control plot on the spit of land between watersheds 2 & 3 just above the parking area, 4. MMS’ Bridge plots (house plot and control plot) near the main trail at EVFS just before the Sonadora footbridge, 5. MMS’ River plots (house plot and control plot) on the lower trail at EVFS near the Sonadora swimming hole, 6. MMS’ Tower 1 plots (House plot and control plot) near the Odum tower and cylinder site, 7. MMS’ Tower 2 plots (house plot and control plot) above the Odum site beside the oxcart path, 8. MMS’ lowsite in the pasture adjacent to Rt 3 near Rio Grande, 9. MMS’ highsite beside the El Toro trail off Rt 191 near the Parrot project. (LLW is Larry Woolbright and MMS is Margaret Stewart)

INVESTIGATORS:

PRINCIPAL INVESTIGATORS

E-MAIL ADDRESS:

Larry Woolbright

lwoolbright@lternet.edu


OTHER RESEARCHERS

E-MAIL ADDRESS:

Margaret Stewart deceased

CONTACT PERSONS

Phone Number (Include area code)

E-MAIL ADDRESS:

Larry Woolbright

(518) 783-2440

lwoolbright@lternet.edu

SOURCE OF FUNDING (SPONSOR):NSF Luquillo LTER; DEB 77-21349 from the National Science Foundation to F. Pough, P. Brussard, and M. Stewart

DATA SET ABSTRACT: We recorded the pattern morph for 9,950 frogs captured at 9 locations in northeastern Puerto Rico over a 25-year period from 1978 - 2002. Data revealed 21 distinct pattern morphs including a variety of stripes, bars, and spots. Analysis of morph frequencies between plots showed significant heterogeneity, with longitudinal stripes more common in grassland and disturbed areas, and spot and bar morphs more common in forests where palm and bromeliad axils are important habitat features. Comparison of morph frequencies through time at the same sites showed temporal shifts immediately following Hurricane Hugo in 1989. We suggest that the pattern polymorphism is maintained in part by local habitat matching resulting from selection pressure from visual predators.

DATA SET METHODS: Plots of varying sizes were searched between 2000 – 2400 h. Adult coquíes were captured by hand and examined to determine their dorsal pattern morphology.
Comparable frequencies between LLW and MMS plots:

LLW M9

MMS M1+M2+M3+M4+M7+M8+M9+M10+M19+M20

LLW M11

MMS M11+M12+M13

LLW M6

MMS M6+M14

LLW M11w

MMS M11a+M15

LLW M16 MMS M16+M17+M18

REFERENCES:
Woolbright, L.L. 2005. A plot-based system of collecting population information on terrestrial breeding frogs. Herpetological Review 36:139-142.

Stewart, M.M. and F.H. Pough. 1983. Population density of tropical forest frogs: relation to retreat sites. Science 221:570-572.

CROSS-REFERENCES (other data sets related to this one): LTERDBAS25: Frog transect data, LTERDBAS30: Frog grid data (Bisley), LTERDBAS44: Frog population plots (El Verde + Bisley), LTERDBAS45: Frog spatial distribution data (El Verde + Bisley), LTERDBAS76: Frog abundance in fertilization plots, LTERDBAS150: Pattern morphology for frogs captured at 9 locations in northeastern Puerto Rico over a 25-year period from 1978 to 2002

SAMPLE LOCATION: N/A

STORAGE SITES(of data files): Originals – LLW, Siena College, Loudonville NY 12211 USA

INVESTIGATOR'S ASSIGNED KEYWORD:
polymorphism, predation, disturbance, camouflage, cryptic, disruptive

LEF LTER OFFICIAL KEYWORDS (See table):LEF, POPULATION DYNAMICS, AMPHIBIANS, PEER REVIEWED JOURNAL

PUBLICATIONS:
Woolbright, L.L. and M.M. Stewart. in press. Spatial and Temporal Variation in Color Pattern Morphology in the Tropical Frog, Eleutherodactylus coqui. Copeia

DISSEMINATION:

RESTRICTED ___ UNRESTRICTED _X__

REASONS TO RESTRICT DATA IN THIS DATA SET BEYOND ITS TWO YEAR POLICY PERIOD*:

*WILL HAVE TO BE APPROVED BY AT LEAST ONE LUQ LTER PRINCIPAL INVESTIGATORS: N. Brokaw, J. ZIMMERMAN, A. LUGO

FILING

_X__ "File" copy only:Data Management will only file an electronic copy of the data file and its documentation 
___ " Enter" data on-line:Data Management will be in charge of entering the data on computer files (Contact Eda C. Meléndez)

SITES DESCRIPTIONS:

Geographical positional system (GPS) Coordinates for each location:

location

latitude

longitude

     

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)

Sonadora, Bisley, S2Bisley, FCMMS plots Sonadora, Bisley, S2Bisley, FCMMS plots Sonadora, Bisley, S2Bisley, FCMMS plots Sonadora, Bisley, S2Bisley, FCMMS plots Sonadora, Bisley, S2Bisley, FCMMS plots

AbbreviationAbbreviation(as it appears on the data file)

season M05 M06 M09 M11

NAME OF VARIABLE

Season period of observation Morph 5 Morph 6 Morph 9 Morph 11

DEFINITION OF VARIABLE

Date or period of time where the observations when done (mm/dd/yyyy) Exact count of number of frogs with white dots on back and heels Exact count of number of frogs with staight interocular band Exact count of number of frogs with mottled with irregular dorsolateral bands Exact count of number of frogs with light vertebral hairline stripe

UNIT

  number number number number

PRECISION

         

RANGE OR LIST OF VALUES

  0,... 0,... 0,... 0,...

DATA TYPE

datetime integer integer integer integer

MISSING DATA CODES

none N/A N/A N/A N/A

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)

Sonadora, Bisley, S2Bisley, FCMMS plots Sonadora, Bisley, S2Bisley, FCMMS plots MMS plots MMS plots MMS plots

AbbreviationAbbreviation(as it appears on the data file)

M11w M16 plot M01 M02

NAME OF VARIABLE

Morph 11 wide Morph 16 Name of plot Morph 1 Morph 2

DEFINITION OF VARIABLE

Exact count of number of frogs with wide vertebral stripe Exact count of number of frogs with light dorsolateral stripes Name given to the survey location Exact count of number of frogs with uniform color throughout Exact count of number of frogs with dark mottled

UNIT

number number   number number

PRECISION

         

RANGE OR LIST OF VALUES

0,... 0,... bridge control, bridge house, river control, river house + grid, tower1 control, tower1 house, tower2 control, tower2 house, lowsite, highsite 0,... 0,...

DATA TYPE

integer integer alphanumeric integer integer

MISSING DATA CODES

N/A N/A none N/A N/A

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)

MMS plots MMS plots MMS plots MMS plots

AbbreviationAbbreviation(as it appears on the data file)

M03 M04 M07 M08

NAME OF VARIABLE

Morph 3 Morph 4 Morph 7 Morph 8

DEFINITION OF VARIABLE

Exact count of number of frogs with vagur interocular bar triangle Exact count of number of frogs with white circles on snout Exact count of number of frogs with dark interocular band; light anteriorly Exact count of number of frogs with vague X on back; light leg stripes

UNIT

number number number number

PRECISION

       

RANGE OR LIST OF VALUES

0,... 0,... 0,... 0,...

DATA TYPE

integer integer integer integer

MISSING DATA CODES

N/A N/A N/A N/A

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)

MMS plots MMS plots MMS plots MMS plots MMS plots

AbbreviationAbbreviation(as it appears on the data file)

M10 M12 M13 M14 M15

NAME OF VARIABLE

Morph 10 Morph 12 Morph 13 Morph 14 Morph 15

DEFINITION OF VARIABLE

Exact count of number of frogs with light sides and snout Exact count of number of frogs with vertebral hairline plus circles on snout Exact count of number of frogs with vertebral hairline plus light snout Exact count of number of frogs with wide vertebral stripe plus interocular bar Exact count of number of frogs with wide vertebral stripe plus circles on snout

UNIT

number number number number number

PRECISION

         

RANGE OR LIST OF VALUES

0,... 0,... 0,... 0,... 0,...

DATA TYPE

integer integer integer integer integer

MISSING DATA CODES

N/A N/A N/A N/A N/A

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)

MMS plots MMS plots MMS plots MMS plots

AbbreviationAbbreviation(as it appears on the data file)

M17 M18 M19 M20

NAME OF VARIABLE

Morph 17 Morph 18 Morph 19 Morph 20

DEFINITION OF VARIABLE

Exact count of number of frogs with dorsolateral stripes ending at eyes Exact count of number of frogs with dorsolateral stripes and light snout Exact count of number of frogs with white upper lip Exact count of number of frogs with white stripes from eyes to nostrils

UNIT

number number number number

PRECISION

       

RANGE OR LIST OF VALUES

0,... 0,... 0,... 0,...

DATA TYPE

integer integer integer integer

MISSING DATA CODES

N/A N/A N/A N/A

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)

MMS plots MMS plots MMS plots MMS plots MMS plots

AbbreviationAbbreviation(as it appears on the data file)

1+9 2+6 2+9 3+8 3+11

NAME OF VARIABLE

Combination Morph 1 and 9 Combination Morph 2 and 6 Combination Morph 2 and 9 Combination Morph 3 and 8 Combination Morph 3 and 11

DEFINITION OF VARIABLE

Exact count of number of frogs with uniform color throughout and mottled with irregular dorsolateral bands Exact count of number of frogs with dark mottled and staight interocular band Exact count of number of frogs with dark mottled and mottled with irregular dorsolateral bands Exact count of number of frogs with vagur interocular bar triangle and vague X on back; light leg stripes Exact count of number of frogs with vagur interocular bar triangle and light vertebral hairline stripe

UNIT

number number number number number

PRECISION

         

RANGE OR LIST OF VALUES

0,... 0,... 0,... 0,... 0,...

DATA TYPE

integer integer integer integer integer

MISSING DATA CODES

N/A N/A N/A N/A N/A

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)

MMS plots MMS plots MMS plots MMS plots MMS plots

AbbreviationAbbreviation(as it appears on the data file)

4+9 4+11w 4+18 5+11 5+16

NAME OF VARIABLE

Combination Morph 4 and 9 Combination Morph 4 and 11w Combination Morph 4 and 18 Combination Morph 5 and 11 Combination Morph 5 and 16

DEFINITION OF VARIABLE

Exact count of number of frogs with white circles on snout and mottled with irregular dorsolateral bands Exact count of number of frogs with white circles on snout and wide vertebral stripe Exact count of number of frogs with white circles on snout and dorsolateral stripes and light snout Exact count of number of frogs with white dots on back and heels and light vertebral hairline stripe Exact count of number of frogs with white dots on back and heels and light dorsolateral stripes

UNIT

number number number number number

PRECISION

         

RANGE OR LIST OF VALUES

0,... 0,... 0,... 0,... 0,...

DATA TYPE

integer integer integer integer integer

MISSING DATA CODES

N/A N/A N/A N/A N/A

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)

MMS plots MMS plots MMS plots MMS plots MMS plots

AbbreviationAbbreviation(as it appears on the data file)

6+9 6+11 6+11w 6+16 6+17

NAME OF VARIABLE

Combination Morph 6 and 9 Combination Morph 6 and 11 Combination Morph 4 and 18 Combination Morph 6 and 16 Combination Morph 6 and 17

DEFINITION OF VARIABLE

Exact count of number of frogs with staight interocular band and mottled with irregular dorsolateral bands Exact count of number of frogs withstaight interocular band and light vertebral hairline stripe Exact count of number of frogs with staight interocular band and wide vertebral stripe Exact count of number of frogs with staight interocular band and light dorsolateral stripes Exact count of number of frogs with staight interocular band and dorsolateral stripes ending at eyes

UNIT

number number number number number

PRECISION

         

RANGE OR LIST OF VALUES

0,... 0,... 0,... 0,... 0,...

DATA TYPE

integer integer integer integer integer

MISSING DATA CODES

N/A N/A N/A N/A N/A

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)

MMS plots MMS plots MMS plots MMS plots MMS plots

AbbreviationAbbreviation(as it appears on the data file)

6+18 7+11w 8+9 8+10 9+10

NAME OF VARIABLE

Combination Morph 6 and 18 Combination Morph 7 and 11w Combination Morph 8 and 9 Combination Morph 8 and 10 Combination Morph 9 and 10

DEFINITION OF VARIABLE

Exact count of number of frogs with staight interocular band and dorsolateral stripes and light snout Exact count of number of frogs with dark interocular band; light anteriorly and wide vertebral stripe Exact count of number of frogs with vague X on back; light leg stripes and mottled with irregular dorsolateral bands Exact count of number of frogs with vague X on back; light leg stripes and light sides and snout Exact count of number of frogs with mottled with irregular dorsolateral bands and light sides and snout

UNIT

number number number number number

PRECISION

         

RANGE OR LIST OF VALUES

0,... 0,... 0,... 0,... 0,...

DATA TYPE

integer integer integer integer integer

MISSING DATA CODES

N/A N/A N/A N/A N/A

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)

MMS plots MMS plots MMS plots MMS plots MMS plots

AbbreviationAbbreviation(as it appears on the data file)

9+11 9+16 9+19 10+13 11+13

NAME OF VARIABLE

Combination Morph 6 and 18 Combination Morph 9 and 16 Combination Morph 9 and 19 Combination Morph 10 and 13 Combination Morph 11 and 13

DEFINITION OF VARIABLE

Exact count of number of frogs with mottled with irregular dorsolateral bands and dorsolateral stripes and light snout Exact count of number of frogs with mottled with irregular dorsolateral bands and light dorsolateral stripes Exact count of number of frogs with mottled with irregular dorsolateral bands and white upper lip Exact count of number of frogs with light sides and snout and vertebral hairline plus light snout Exact count of number of frogs with mottled with light vertebral hairline stripe and vertebral hairline plus light snout

UNIT

number number number number number

PRECISION

         

RANGE OR LIST OF VALUES

0,... 0,... 0,... 0,... 0,...

DATA TYPE

integer integer integer integer integer

MISSING DATA CODES

N/A N/A N/A N/A N/A

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)

MMS plots MMS plots MMS plots MMS plots MMS plots

AbbreviationAbbreviation(as it appears on the data file)

11+16 11+17 11+19 11+20 11w+7

NAME OF VARIABLE

Combination Morph 11 and 16 Combination Morph 11 and 17 Combination Morph 11 and 19 Combination Morph 11 and 20 Combination Morph 11w and 7

DEFINITION OF VARIABLE

Exact count of number of frogs with with light vertebral hairline stripe and light dorsolateral stripes Exact count of number of frogs with with light vertebral hairline stripe and dorsolateral stripes ending at eyes Exact count of number of frogs with with light vertebral hairline stripe and white upper lip Exact count of number of frogs with with light vertebral hairline stripe and white stripes from eyes to nostrils Exact count of number of frogs with wide vertebral stripe and dark interocular band; light anteriorly

UNIT

number number number number number

PRECISION

         

RANGE OR LIST OF VALUES

0,... 0,... 0,... 0,... 0,...

DATA TYPE

integer integer integer integer integer

MISSING DATA CODES

N/A N/A N/A N/A N/A

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)

MMS plots MMS plots MMS plots MMS plots MMS plots

AbbreviationAbbreviation(as it appears on the data file)

11w+16 13+16 13+18 13+20 15+16

NAME OF VARIABLE

Combination Morph 11 and 16 Combination Morph 13 and 16 Combination Morph 13 and 18 Combination Morph 13 and 20 Combination Morph 15 and 16

DEFINITION OF VARIABLE

Exact count of number of frogs with wide vertebral stripe and light dorsolateral stripes Exact count of number of frogs with vertebral hairline plus light snout and wide and light dorsolateral stripes Exact count of number of frogs with vertebral hairline plus light snout and dorsolateral stripes and light snout Exact count of number of frogs with vertebral hairline plus light snout and white stripes from eyes to nostrils Exact count of number of frogs with wide vertebral stripe plus circles on snout and light dorsolateral stripes

UNIT

number number number number number

PRECISION

         

RANGE OR LIST OF VALUES

0,... 0,... 0,... 0,... 0,...

DATA TYPE

integer integer integer integer integer

MISSING DATA CODES

N/A N/A N/A N/A N/A

VARIABLES (ATTRIBUTES):

File Name or # above (all in which the variable appears)

MMS plots MMS plots

AbbreviationAbbreviation(as it appears on the data file)

16+18 17+19

NAME OF VARIABLE

Combination Morph 16 and 18 Combination Morph 17 and 19

DEFINITION OF VARIABLE

Exact count of number of frogs with light dorsolateral stripes and light snout Exact count of number of frogs with dorsolateral stripes ending at eyes and white upper lip

UNIT

number number

PRECISION

   

RANGE OR LIST OF VALUES

0,... 0,...

DATA TYPE

integer integer

MISSING DATA CODES

N/A N/A

COMPUTATIONAL METHODS:

Variable Name

Formula

   

 



FOR DATA MANAGER USE ONLY

DATE OF LAST REVIEW: July 9, 2007

DATE OF LAST ENTRY: August 2, 2002

STAGE OF DATA SET MANAGEMENT (dates):
RECEIVED: March 26, 2007 (data set abstract only)
ENTERED: March 28, 2007; all: metadata and datain June 8, 2007
FILED ON-LINE:
REVIEWED BY RESEARCHER: Jul 02, 2007

FILING MEDIA:
NAME OF DOCUMENTATION FILE:
lterdb150.htm
NAME OF ON - LINE CATALOG:
LTERDBAS
RECORD #:
150
DOCUMENT TYPE:
electronic media only
PRIORITY TO BE ENTERED: N/A

Rev. date of this form: 28 July 200/ 15 July  2001/June 9, 2003/March 16, 2004/12 April 2005/ 8 November 2005/ 16 January 2006