LUQ LTER DATA SETS DOCUMENTATION FORM

ON-LINE VERSION

A DATA SET is a series of observations collected by the same methodology. Each data set should have documentation sufficient for someone unfamiliar with the research to replicate the study. Data sets may be broken into subsets (data files) that are discrete in space and time, in that order. The documentation for a data set should include all spatial and temporal subdivisions of the data.

(Data, Abstract, Methods, Variables)

NOTES:

PERSON(S) COMPLETING THIS FORM: E-MAIL ADDRESS:

Christopher P. Bloch

cbloch@ttacs.ttu.edu

DATA SET IDENTIFIER: Bisley Grid Invertebrate Data, 1989-1999

PROJECT TITLE: Community Ecology of Land Snails (Long-term population dynamics of snails in the Tabonuco forest)

PROJECT DESCRIPTION: Terrestrial gastropods and walking sticks were sampled at 36-40 points on the grid at the Bisley Watersheds to investigate long-term population and community dynamics and response to disturbance.

Densities

Densities of invertebrates and apparency of plant species are estimated to investigate the effects of disturbance on long-term population dynamics, habitat associations, and spatial distributions.

Habitat

To investigate habitat associations of terrestrial invertebrates, apparency of 17 plant groups (species, genera, families) from 0-3 m from the ground was estimated at 36 points on the grid at Bisley Watersheds 1 and 2 in 1994 and 1999.

LTER CORE AREAS: (Annotate all that apply)
Population Dyanamics
Disturbance Patterns

LEF LTER 1 RESEARCH TOPIC: (Annotate all that apply)
Recovery After Disturbance
Spatial and Temporal Distribution of Populations

We define a data file as a component of a data set. A data set can have only one data file or more. Basically, different data files have different data structures or format.
DATA SET FILES (SUBSETS):

Data File No.

Data File Identifier

On-Line Filename

Starting Date

Periodicity of sample

End Period

1 Caracolus caracolla abundance 1989-1999 Cc.txt

June 1, 1989

Irregular (1989,1990,1994,1999)
August 7, 1999
2 Gaeotis nigrolineata abundance 1989-1999 Gn.txt June 1, 1989
Irregular (1989,1990,1994,1999)
August 7, 1999
3 Nenia tridens abundance 1989-1999 Nt.txt June 1, 1989
Irregular (1989,1990,1994,1999)
August 7, 1999
4 Polydontes acutangula abundance 1989-1999 Pa.txt June 1, 1989
Irregular (1989,1990,1994,1999)
August 7, 1999
5 Abundance of all snail species and walking sticks in 1994 Bisleydatasummary1994.txt

June 20, 1994

3 times, roughly bi-weekly
July 7, 1994
6 Abundance of all snail species and walking sticks in 1999 Bisleydatasummary1999.txt August, 1999
Single sample of each quadrat over approx. 1 week
August 7, 1999

RESEARCH LOCATION: Bisley grid

INVESTIGATORS:

PRINCIPAL INVESTIGATORS E-MAIL address

Michael R. Willig

michael.willig@ttu.edu
mwillig@nsf.gov

OTHER RESEARCHERS E-MAIL address
Christopher Bloch cbloch@ttacs.ttu.edu
Gerardo Camilo camilogr@sluava.slu.edu
Michelle Wolf mwolf@montana.edu

CONTACT PERSONS E-MAIL address Phone Number (Include area code)

M. Willig or C. Bloch

michael.willig@ttu.edu
mwillig@nsf.gov

(806)742-2658 (Texas Tech Univ.), or (703)292-7198 (Willig at NSF)


SOURCE OF FUNDING (SPONSOR): NSF (Luquillo LTER), U.S. Dept. of Energy (Oak Ridge Associated Universities)

DATA SET ABSTRACT: The data set consists of counts of terrestrial invertebrates from the grid at Bisley Watersheds #1 and 2, for the years 1989, 1990, 1994, and 1999. Data for 1989 and 1990 are confined to 4 species of terrestrial snails: Caracolus caracolla, Gaeotis nigrolineata, Nenia tridens, and Polydontes acutangula. Counts for other snail species and the walking stick Lamponius portoricensis are available for 1994 and 1999.

DATA SET METHODS: Thirty six points were selected from the grid at Bisley based upon accessibility during night-time surveys (for a map of the study plots, see Willig and Camilo, 1991). See the List of Codes of the 36 selected grid points. Circular quadrats (r = 5 m) were established at each point and sampled during the wet season immediately prior to Hurricane Hugo (June-August 1989), that immediately following Hugo (July-August 1990), and in July-August of 1994 and August of 1999. Sampling at each of the selected points consisted of a minimum of two individuals searching all available surfaces within a quadrat (e.g., vegetation, litter, rock, soil) for the presence of snails. All surveys occurred between 19:30 and 03:00 hours to coincide with peak snail activity.

REFERENCES:

Willig, M.R., and G.R. Camilo. 1991. The effect of Hurricane Hugo on six invertebrate species in the Luquillo
Experimental Forest of Puerto Rico. Biotropica 23:455-461.

CROSS-REFERENCES (other data sets related to this one): LTERDBAS 42: Grid Habitat Data, Bisley 1994,1999

SAMPLE LOCATION: N/A

STORAGE SITES (of data files): Data Management Files DM-003 Drawer 4

INVESTIGATOR'S ASSIGNED KEYWORDS: snails, walking sticks, population density, disturbance, Hurricane Hugo

LEF LTER OFFICIAL KEYWORDS (See table): BISLEY WATERSHEDS: Q1, Q2, DISTURBANCE, HURRICANE, MOLLUSKS, INSECTS, PEER REVIEWED JOURNAL

PUBLICATIONS:
Secrest, M.F. 1995. The impact of Hurricane Hugo on two common tree snails in the Luquillo Experimental Forest
of Puerto Rico: a long-term study. M.S. Thesis, Texas Tech University, Lubbock, Texas.

Secrest, M.F., M.R. Willig, and L.L. Peppers. 1996. The legacy of disturbance on habitat associations of terrestrial
snails in the Luquillo Experimental Forest, Puerto Rico. Biotropica 28:502-514.

Willig, M.R., and G.R. Camilo. 1991. The effect of Hurricane Hugo on six invertebrate species in the Luquillo
Experimental Forest of Puerto Rico. Biotropica 23:455-461.

Willig, M.R., M.F. Secrest, J.F. Cary, S.B. Cox, G.R. Camilo, and M.R. Gannon. 1998. Long-term monitoring of snails in the Luquillo Experimental Forest of Puerto Rico: heterogeneity, scale, disturbance, and recovery. In F. Dallmeier and J. Comisky, editors. Forest Biodiversity in North, Central, and South America and the Caribbean: Research and Monitoring. UNESCO and The Parthenon Press, Carnforth, Lancashire, UK. Pp. 293-322.


DISSEMINATION:
UNRESTRICTED

REASONS TO RESTRICT DATA IN THIS DATA SET BEYOND ITS TWO YEAR POLICY PERIOD*:

*WILL HAVE TO BE APPROVED BY LTER PRINCIPAL INVESTIGATORS: J. ZIMMERMAN, A. LUGO , D.J. LODGE

SITES DESCRIPTIONS: This study was conducted in two of the Bisley Experimental Watersheds of the Luquillo Experimental Forest (LEF) of Puerto Rico. The Luquillo mountains (18° 20'N, 65° 49'W) are the predominant physiographic feature in northeastern Puerto Rico and are from sea level to an elevation of 1075 m over a distance of approximately 10 km. The Bisley watersheds 1 and 2 are on the windward side of the mountains, have a mean annual precipitation of approximately 3500 mm, and occur in the subtropical forest life zone (Scatena, 1989). These adjacent watersheds drain a total of 13 highly dissected hectares that are underlain by a Cretaceous age volcanoclastic oldstone of andesitic composition.

Geographical positional system (GPS) Coordinates for each location:

location

latitude

longitude

Bisley Watershed 1    
Bisley Watershed 2    

VARIABLES (ATTRIBUTES):

1FILE NAME OR #ABOVE (all in which the variable appears)

All All 1 1 1 1

ABBREVIATION (as it appears on the data file)

X axis Y Axis CC_89 CC_90 CC_94 CC_99

NAME OF VARIABLE

X coordinate of the point on the grid observed Y coordinate of the point on the grid observed Abundance of Caracolus caracolla in 1989 Abundance of Caracolus caracolla in 1990 Abundance of Caracolus caracolla in 1994 Abundance of Caracolus caracolla in 1999

DEFINITION OF VARIABLE

X coordinate of the point on the grid (quadrat coordinate where the observations where performed) Y coordinate of the point on the grid (quadrat coordinate where the observations where performed) Minimum Number Known Alive of Caracolus caracolla individuals when observations where done in 1989 Minimum Number Known Alive of Caracolus caracolla individuals when observations where done in 1990 Minimum Number Known Alive of Caracolus caracolla individuals when observations where done in 1994 Minimum Number Known Alive of Caracolus caracolla individuals when observations where done in 1999

UNIT

number number number number

number

 

number

 

PRECISION

           

RANGE OR LIST OF VALUES

           

DATA TYPE

 integer  integer integer integer integer integer

MISSING DATA CODES

    none none none none

VARIABLES (ATTRIBUTES):

2FILE NAME OR #ABOVE (all in which the variable appears)

2 2 2 2

ABBREVIATION (as it appears on the data file)

GN_89 GN_90 GN_94 GN_99

NAME OF VARIABLE

Abundance of Gaeotis nigrolineata in 1989 Abundance of Gaeotis nigrolineata in 1990 Abundance of Gaeotis nigrolineata in 1994 Abundance of Gaeotis nigrolineata in 1999

DEFINITION OF VARIABLE

Minimum Number Known Alive of Gaeotis nigrolineata individuals when observations where done in 1989 Minimum Number Known Alive of Gaeotis nigrolineata individuals when observations where done in 1990 Minimum Number Known Alive of Gaeotis nigrolineata individuals when observations where done in 1994 Minimum Number Known Alive of Gaeotis nigrolineata individuals when observations where done in 1999

UNIT

number

 

number

 

number

 

number

 

PRECISION

       

RANGE OR LIST OF VALUES

       

DATA TYPE

integer integer integer integer

MISSING DATA CODES

none none none none

VARIABLES (ATTRIBUTES):

3FILE NAME OR #ABOVE (all in which the variable appears)

3 3 3 3

ABBREVIATION (as it appears on the data file)

NT_89 NT_90 NT_94 NT_99

NAME OF VARIABLE

Abundance of Nenia tridens in 1989 Abundance of Nenia tridens in 1990 Abundance of Nenia tridens in 1994 Abundance of Nenia tridens in 1999

DEFINITION OF VARIABLE

Minimum Number Known Alive of Nenia tridens individuals when observations where done in 1989 Minimum Number Known Alive of Nenia tridens individuals when observations where done in 1990 Minimum Number Known Alive of Nenia tridens individuals when observations where done in 1994 Minimum Number Known Alive of Nenia tridens individuals when observations where done in 1999

UNIT

number

 

number

 

number

 

number

 

PRECISION

       

RANGE OR LIST OF VALUES

       

DATA TYPE

integer integer integer integer

MISSING DATA CODES

none none none none

VARIABLES (ATTRIBUTES):

4FILE NAME OR #ABOVE (all in which the variable appears)

4 4 4 4

ABBREVIATION (as it appears on the data file)

PA_89 PA_90 PA_94 PA_99

NAME OF VARIABLE

Abundance of Polydontes acutangula in 1989 Abundance of Polydontes acutangula in 1990 Abundance of Polydontes acutangula in 1994 Abundance of Polydontes acutangula in 1999

DEFINITION OF VARIABLE

Minimum Number Known Alive of Polydontes acutangula individuals when observations where done in 1989 Minimum Number Known Alive of Polydontes acutangula individuals when observations where done in 1990 Minimum Number Known Alive of Polydontes acutangula individuals when observations where done in 1994 Minimum Number Known Alive of Polydontes acutangula individuals when observations where done in 1999

UNIT

number

 

number

 

number

 

number

 

PRECISION

       

RANGE OR LIST OF VALUES

       

DATA TYPE

integer integer integer integer

MISSING DATA CODES

none none none none

VARIABLES (ATTRIBUTES):

5FILE NAME OR #ABOVE (all in which the variable appears)

5,6 5,6 5,6 5,6 5,6

ABBREVIATION (as it appears on the data file)

Season Date Aa As Cc

NAME OF VARIABLE

Season code Date of observation Abundance of Alcadia alta Abundance of Alcadia striata Abundance of Caracolus caracolla

DEFINITION OF VARIABLE

Season code containing the last two digit of the year, the season (W for wet), and in the summary of 1999 a digit representing the date in which the observation was done (1 for 06/20/94, 2 for 07/04/94, and 3 for 07/24/94) Date when observation was done Total of Minimum Number Known Alive of Alcadia alta observed individuals observed in the plot in the corresponding date Total number of Minimum Number Known Alive of Alcadia striata individuals observed in the plot in the corresponding date Total number of Minimum Number Known Alive of Caracolus caracolla individuals observed in the plot in the corresponding date

UNIT

    number number number

PRECISION

         

RANGE OR LIST OF VALUES

94W1, 94W2, 94W3 06/20/94
07/04/94
07/24/94
08/01/99
     

DATA TYPE

alphanumeric date integer integer integer

MISSING DATA CODES

none none none none none

VARIABLES (ATTRIBUTES):

6FILE NAME OR #ABOVE (all in which the variable appears)

5,6 5,6 5,6 6 5,6

ABBREVIATION (as it appears on the data file)

Cm Cs Gn Lg Mc

NAME OF VARIABLE

Abundance of Caracolus marginella Abundance of Cepolis squamosa Abundance of Gaeotis nigrolineata Abundance of Lamellaxis gracilis Abundance of Megalomastoma croceum

DEFINITION OF VARIABLE

Total of Minimum Number Known Alive of Caracolus marginella individuals observed in the plot in the corresponding date Total of Minimum Number Known Alive of Cepolis squamosa individuals observed in the plot in the corresponding date Total of Minimum Number Known Alive of Gaeotis nigrolineataindividuals observed in the plot in the corresponding date Total of Minimum Number Known Alive of Lamellaxis gracilis individuals observed in the plot in the corresponding date Total of Minimum Number Known Alive of Megalomastoma croceum individuals observed in the plot in the corresponding date

UNIT

number number number number number

PRECISION

         

RANGE OR LIST OF VALUES

         

DATA TYPE

integer integer integer integer integer

MISSING DATA CODES

none none none none none

VARIABLES (ATTRIBUTES):

7FILE NAME OR #ABOVE (all in which the variable appears)

5,6 5,6 5,6 6 5,6

ABBREVIATION (as it appears on the data file)

Nt Og Op Ot Pa

NAME OF VARIABLE

Abundance of Nenia tridens Abundance of Oleacina glabra Abundance of Oleacina playa Abundance of Obeliscus terebraster Abundance of Polydontes acutangula

DEFINITION OF VARIABLE

Total of Minimum Number Known Alive of Nenia tridens individuals observed in the plot in the corresponding date Total of Minimum Number Known Alive of Oleacina glabra individuals observed in the plot in the corresponding date Total of Minimum Number Known Alive of Oleacina playa individuals observed in the plot in the corresponding date Total of Minimum Number Known Alive of Obeliscus terebraster individuals observed in the plot in the corresponding date Total of Minimum Number Known Alive of Polydontes acutangul individuals observed in the plot in the corresponding date

UNIT

number number number number number

PRECISION

         

RANGE OR LIST OF VALUES

         

DATA TYPE

integer integer integer integer integer

MISSING DATA CODES

none none none none none

VARIABLES (ATTRIBUTES):

8FILE NAME OR #ABOVE (all in which the variable appears)

5,6 5,6 5,6 5,6 6

ABBREVIATION (as it appears on the data file)

Pp So Vo Total Snails Lp

NAME OF VARIABLE

Abundance of Platysuccinea portoricensis Abundance of Subulina octana Abundance of Vaginulus occidentalis Total bundance of snails Abundance of Lamponius portoricensis, a walking stick

DEFINITION OF VARIABLE

Total of Minimum Number Known Alive of Platysuccinea portoricensis individuals observed in the plot in the corresponding date Total of Minimum Number Known Alive of Subulina octana individuals observed in the plot in the corresponding date Total of Minimum Number Known Alive of Vaginulus occidentalis individuals observed in the plot in the corresponding date Sum of all the M inimum Number Known Alive of snails of all species found at a point Total number of Minimum Number Known Alive walking stick individuals, Lamponius portoricensis. Juveniles are denoted by "j".

UNIT

number number number number number

PRECISION

         

RANGE OR LIST OF VALUES

         

DATA TYPE

integer integer integer integer integer

MISSING DATA CODES

none none none none none

COMPUTATIONAL METHODS:
Variable Name Formula
   


FOR DATA MANAGER USE ONLY

DATE OF LAST REVIEW: February 16, 2008
DATE OF LAST ENTRY:

STAGE OF DATA SET MANAGEMENT (dates):

RECEIVED Sep 29, 2000

ENTERED: Oct 4, 2000

FILED ON-LINE Oct 4, 2000

REVIEWED BY RESEARCHER

FILING MEDIA:

NAME OF DOCUMENTATION FILE: lterdb43.htm,  BisleyGrid.doc

NAME OF ON - LINE CATALOG: LTERDBAS    RECORD # :43

DOCUMENT TYPE:

PRIORITY TO BE ENTERED : N/A


Rev. date of this form: 6 March 2002