2 File Name or # above (all in which the variable appears)

Abbreviation (as it appears on the data file)

NAME OF VARIABLE

DEFINITION OF VARIABLE

UNIT

PRECISION

RANGE OR LIST OF VALUES

DATA TYPE

MISSING DATA CODES

2 File Name or # above (all in which the variable appears)

13

13

13

1-15, 20

Abbreviation (as it appears on the data file)

CUTMS

CUTCODE

CUTPET

DDIF

NAME OF VARIABLE

Cut cycle month

Cut code

Cut petiole

Day difference

DEFINITION OF VARIABLE

Number of months from the beginning of the study, September, 1991, to the month of monitoring period in which leaf was obviously cut by machete in clearing trails.

Code indicating that leaf has been cut in clearing trails.

Number of pairs of leaflets left after leaf is cut. If blank, only petiole remained.

Difference between date first observed and date next observed during a single monitoring period.

UNIT

PRECISION

RANGE OR LIST OF VALUES

{1,.,99}

C = Leaf cut by machete, U = Plant bulldozed

1-31

DATA TYPE

numeric

alphanumeric

alphanumeric

numeric

MISSING DATA CODES

Present only if leaf has been cut

Blank if not applicable

Blank if not applicable.

Must be present when PR code is D or E.

3 File Name or # above (all in which the variable appears)

1-15, 18, 20

1-15, 20

1-17, 20

1-15, 20

Abbreviation (as it appears on the data file)

DAY

DIF

GPHEN

GC

NAME OF VARIABLE

Day re-monitored

Difference in length

Leaf or plant gone season code

Gone code

DEFINITION OF VARIABLE

Date of month that first and subsequent measurements were taken during the same monitoring month.

Difference between: 1) PR code = R: leaf length and length on P record (if not first B record) or previous R record if there is one and Status of leaf is a B or blank if diff. is on a record with PR code P. 2) PR code = S: leaf length and length on P or R

The year is divided into three "seasons" (between monitoring dates). Leaf or plant gone season is the season in which the leaf or plant was assumed to have died as noted at the next monitoring date early in the month following the end of the season. If annual monitoring, then 0 indicates leaf or plant gone during the previous year. On leaf monitoring files, is found only the records with PR code of P.

Code indicating the condition under which a leaf was no longer present in the study.

UNIT

centimeter

PRECISION

RANGE OR LIST OF VALUES

1-31

-999 to 999

0 = January/December, 1 = January/April, 2 = May/August, 3 = September/December

Leaf phenology/life span codes : G = no sign of leaf after four months, K = Same as G but plant not observed previous visit (after 8 months), N = same as G but plant not seen for two visits prior to this one (12 months) or since the last annual monitoring visit (Phase 2). All leaf records with MYRPHEN greater then 2000 (Phase II annual measurements) use this code rather than G., H = wHip-petiole and rachis only present (no leaflets if that is typical of species). Dead with leaflets still attached if that is typical of species (After four months), L = Same as H, but plant not observed previous visit (after 8 months), O = Same as H but plant not seen for two visits prior to this one (12 months) or since the last annual monitoring visit (Phase 2). All leaf records with MYRPHEN greater then 2000 (Phase II annual measurements) use this code rather than H., D = Dead with leaflets still attached if losing leaflets is typical of species, Z = Leaf gone because its plant is assumed or known to be dead (may not be verified until next monitoring period)

Plant (therefore leaf) droppedfrom study codes X = Leaf no longer monitored because plant isn't, though plant is not known to be dead, just not found or dropped for other reasons, thus this is NOT a phen code. If a plant is not measured temporarily but is found later, life span is really not known, but plant found later (special case ofcode X), F = Leaf experimentally defoliated, thus natural life span not known (special case of code X), E = observation of this leaf ended at the end of phase 1, thus its life span is not known (special case of code X).Either the site was dropped (5, 10, Jobos Bay), or the plant went into phase II with some (but not all) of the leaves previously noted so that a specific leaf's life span could not be determined (Thelypteris angustifolia).

Special codes: B = (S or D Profile code): broken by next monitoring period, Q = (C Type code): still immature at next monitoring period, thus unrolling takes more than one season, R = Leaves measured when new Phase II plants were profiled.

DATA TYPE

integer

integer

integer

alphanumeric

MISSING DATA CODES

Must be present when PR code is S, D and/or E if leaf remonitored during the same month.

Blank if not applicable.

Blank if year and/or season of death is not known.

Blank if leaf is present

4 File Name or # above (all in which the variable appears)

1-17, 20

1-17, 20

1-22

1-15, 20

Abbreviation (as it appears on the data file)

GS

GYRPHEN

ID

LAM

NAME OF VARIABLE

Gone cycle month

Gone year

ID number of plant

Lamina length

DEFINITION OF VARIABLE

Number of months from beginning of study (September, 1991) to month of monitoring date when a previously monitored leaf or plant is no longer part of the study. On leaf monitoring records, used only on records with PR code of P. Phase 1, generally every four months (1-113); Phase 2, generally every 12 months (125-240).

Year in which a leaf or plant is observed or assumed to be dead at the next monitoring date.On leaf monitoring records, used only on records with PR code of P, and not on records with a Gone code (GC) of X, T, E, F, Q or B.

The sequentially assigned number on the tag that is affixed to the plant when it is old enough not to be damaged by a tag. For Phase 2, note that ID's greater than 2999and less than 9000 are temporary, do not involve tagging a specific plant and are assigned by year (ex 4000 = 2004)

Length of leaf from start of lamina or lowest pair of leaflets to tip.

UNIT

centimeter

PRECISION

RANGE OR LIST OF VALUES

1-99

1991-2011

1-99999

1-999

DATA TYPE

integer

integer

integer

numeric

MISSING DATA CODES

Blank unless leaf is gone

Blank if leaf is still present or if gone, then year of death not known.

Must be present

Blank if simple leaf or crozier which has not reached the lowest set of leaflets in the unrolling process, or a leaf that broke off in the petiole

5 File Name or # above (all in which the variable appears)

13

13

13

13

Abbreviation (as it appears on the data file)

LAM1I

LAM2I

LAM1C

LAM2C

NAME OF VARIABLE

Lamina 1/Inches

Lamina 2/Inches

Lamina 1/centimeter

Lamina 2/centimeter

DEFINITION OF VARIABLE

On a fertile leaf, the measurement in inches of the area of sterile leaflets along the rachis above the petiole and below the start of the fertile leaflets.

On a fertile leaf, the measurement in inches of the area of fertile leaflets along the rachis above the sterile leaflets.

On a fertile leaf, the measurement in centimeter of the area of sterile leaflets along the rachis above the petiole and below the start of the fertile leaflets. (*Calculated: Lamina 1/Inches * 2.54)

On a fertile leaf, the measurement in centimeter of the area of fertile leaflets along the rachis above the sterile leaflets. (*Calculated: Lamina 2/Inches * 2.54))

UNIT

inches

inches

centimeter

centimeter

PRECISION

RANGE OR LIST OF VALUES

DATA TYPE

numeric

numeric

integer

integer

MISSING DATA CODES

6 File Name or # above (all in which the variable appears)

1-15, 20

1-15, 20

1-15, 20

1-18, 20

Abbreviation (as it appears on the data file)

LEAF

LN

LEN

MS

NAME OF VARIABLE

Leaf condition

Leaf condition modifier

Length of leaf

Monitor cycle month

DEFINITION OF VARIABLE

Code indicates if all of the leaf is present in terms of expected number of leaflets for pinnate leaves and for simple leaves, if a portion of the lamina is missing.

Additional information about the development or damage status of the leaf.

Combined length of petiole and lamina. (Note that for Profile code D final length is noted in Orientation field.) (*Calculated: Sum of petiole and lamina length)

Number of months since the beginning of the study with September, 1991 = 1. Records month in which monitoring took place and is used with GS to calculate leaf life span.

UNIT

N/A

N/A

centimeter

N/A

PRECISION

N/A

N/A

N/A

RANGE OR LIST OF VALUES

A = All there, N = Some not there (was M in earlier field notes), n = number of pairs of leaflets present (if minority), T = terminal leaflet only left

n = missing n leaflets

1-999

1-240; Phase 1, generally every four months (1-113); Phase 2, generally every 12 months (125-240).

DATA TYPE

alphanumeric

alphanumeric or integer

integer

integer

MISSING DATA CODES

Must be present

Present if Leaf condition code present unless too many to count

Must be present

Must be present

7 File Name or # above (all in which the variable appears)

1-23

1-23

1-15, 20

1-15, 20

Abbreviation (as it appears on the data file)

MYRPHEN

MPHEN

OR

PRS

NAME OF VARIABLE

Monitoring period year

Leaf or plant monitoring season code

Orientation of leaf

Pairs of pinnae (leaflets)

DEFINITION OF VARIABLE

Year that a plant was added to the study or a leaf was known to have emerged and/or matured or was remonitored. Indicates the year during which the recorded growth took place, not monitoring date year (ie: if monitored in January)

The year is divided into three "seasons" (between monitoring dates).1) Leaf monitoring: For a leaf with an ADD code, the monitoring season is the season in which the leaf is assumed to have emerged or matured or was remonitored as noted at the next monitoring date early in the month following the end of the season. If there is no ADD code, then MPHEN represents the first time a leaf is measured when a plant is added to ths tudy. 2) Plant monitoring: MPHEN represents the season that the plant observations cover.

Orientation of the leaf as it emerges from the apex with respect to some fixed background feature. Used in combination with wire color to identify a leaf when the plant is monitored.

Number of pairs of leaflets with a clearly defined (though possible very short) petiolule on pinnate leaves. For simple leaves, this field may be either blank or 0. 1. Counted in field, usually on the right side with adaxial surface up. If pairs on both side are counted (occasionally done on smaller leaves) and are unequal in number, then .5 is used to average the number on both sides of rachis. 2. For leaves first encountered as croziers (Profile codes S and D), the number of pairs counted when the leaf is mature is inserted in this field later.

UNIT

N/A

N/A

degree

PRECISION

N/A

N/A

RANGE OR LIST OF VALUES

1991-2011

0 = January/December, 1 = January/April, 2 = May/August, 3 = September/December

n = number of degrees around apex with 0 equal to up; 180 equal to down; 90 equal to left and 270 equal to right when looking at direction of the extension of a leaf. Usually oriented with respect to a noted background feature: uphill upstream or upridge equal to 0 degrees, U equal to uphill, D equal to downhill, L equal to left, R equal to right (facing downhill) were used for CYBO before degree system instituted; For Profile code equal to D; this field is used for the final length of the leaf (cm) and is blank if leaf is broken or gone before measured

1-99

DATA TYPE

integer

integer

alphanumeric

integer

MISSING DATA CODES

Must be present

Must be present

Blank before degree system instituted, after that, should be present

Must be present for pinnate leaves. Blank when first measured if crozier has not completely unrolled. Blank when data analyzed if leaf broken or gone before pairs could be counted.

8 File Name or # above (all in which the variable appears)

13

13

1-15, 20

13

Abbreviation (as it appears on the data file)

PRS1

PRS2

PET

PETI

NAME OF VARIABLE

Pairs 1

Pairs 2

Petiole length

Petiole/Inches

DEFINITION OF VARIABLE

For ACRDAN, on a fertile leaf, the number of pairs of sterile leaflets above the petiole and below the fertile leaflets.

On a fertile leaf of ACRDAN, the number of pairs of fertile leaflets above the sterile leaflets on the rachis

Petiole measured from rhizome to base of lamina or lowest pair of leaflets.(*Calculated: For Acrostichum, the petiole length is calculated (PETI * 2.54))

Length of petiole in inches

UNIT

centimeter

Inches

PRECISION

RANGE OR LIST OF VALUES

1-999

DATA TYPE

integer

integer

integer

integer

MISSING DATA CODES

Blank if petiole indistinguishable from the lamina.

9 File Name or # above (all in which the variable appears)

1-15, 20

1-15, 20

1-22

17

Abbreviation (as it appears on the data file)

PR

SEQ

SITE

SP

NAME OF VARIABLE

Profile code

Sequence of leaf

Site Code

Species code

DEFINITION OF VARIABLE

Code indicating type of monitoring done on the leaf.

The oldest appearing leaf when the plant is first profiled is assigned the sequence number 1. Each younger leaf, and each leaf that emerges during later monitoring periods is assigned the next number in sequence.

Numerical code assigned to area of the forest where monitored ferns are located.

Code for scientific name of fern using standard six digit abbreviation of LUQ LTER

UNIT

PRECISION

RANGE OR LIST OF VALUES

P = profile record, one for each leaf node observed (generally signifies a mature leaf), S = starter record, for leaves which are not mature (type C or I) when first encountered (changed to a P in the next monitoring period if the leaf is gone), D = remeasurement of an immature leaf during the same monitoring period as a Starter measurement was taken; Note (985): if a leaf has matured at the time of re-measurement this code becomes a P (A), though the day remonitored and differences remains, R = remeasurement of a leaf in a monitoring period after the original profile has been done

1-99
(Note: Originally letters of the alphabet were used to indicate sequence of leaf emergence.
As of January, 1994 all letter were converted to numbers: A-1, B-2, C-3, D-4, E-5, F-6, G-7, H-8, I-9, J-10 K-11, L-12, M-13, N-14, O-15, P-16, Q-17, R-18, S-19, T-20 U-21, V-22, W-23, X-24, Y-25, Z-26, AA-27, AB-28, AC-29 AD-30, ETC.)

1 = Sonadora forest below pool near station (DAE1, DANO), 2 = Sonadora pool edge near station (NERI, ELSI, THAN), 3 = Sonadora rocks above old bridge-station side (THAN, THDE, THRE, BLOC), 4 = Across Sonadora bridge, opposite a MRCE plot (NERI) plus (Phase 2) plants along trail downhill of this site to Sonadora (ASSE, ASCU, ELRI, POCR), 5 = Espiritu Santo Tributary-across bridge and uphill to the right-downstream of stream crossing (DAE2, THRE, ELCR, THDE, CYBO), 6 = Across Sonadora below pool near station (1 HEMA), 7 = Hurricane Recovery Plot 62--stream edge (1 DAE1), 8 = Sonadora-hillside before bridge on station side, left of trail to bridge (CYBO), 9 = Intermittent stream bed across trail from 8 (DANO), 10 Espiritu Santo Tributary-across bridge and uphill to right-upstream of stream crossing (ELCR, CYBO, THDE), 11 = Sonadora rocks above old bridge-station side, 5 m downstream from 3 (THAN, BLOC), 12 = Sonadora hillside upstream from 3 (CYBO), 13 = Sonadora forest before bridge on station side, about 40 m toward river from 9 (THRE, DAE1, LILA), 14 = Lars Landslide very near station (BONI), 15 = Sonadora ridgetop above bridge on station side (Everham and Weaver plots in area) (THDE), 16 = Not used, 17 = Largest intermittent stream crossed by trail into forest from station--upstream of log by trail. (HEMA, DANO TETR, BLOC, THRE), 18 = Just before Sonadora bridge to right of trail above metal grating (THRE, THOE, DANO), 19 = Pico del Este in area of MRCE plots, 20 = Walk-up tower at El Verde (one HEMA marked), 21 = Bisley Watershed 2 just upstream from gaging station (DANO marked within 2-3 m of stream bed), 22 = Bisley Watershed 3 just upstream from gaging station(DANO) marked within 2-3 m of stream bed), 23 = Bisley Watershed 4 at trail crossing (THAN), 24 = Bisley Watershed 2 about 30 M upslope toward Watershed 3 (DANO), 25 = Experimental site near Site 8 where THDE and DANO have been defoliated, 26 = Comparative site at Jobos Bay-Sugar Mill Interior (ACDA), 27 = Comparative site at Jobos Bay-Sugar Mill Edge (ACDA), 28 = Comparative site at Jobos Bay-Thermoelectric Plant Open (ACDA), 29 = Comparative site at Jobos Bay-Thermoelectric Plant Edge ACDA, 30 = Just S of Big Grid next to Quadrat 0301 (NERI), 33 = Just into the forest from field station on right (THRE), 34 = Experimental site down hill from Site 1 where additional defoliations were done (CYBO, DAE1), 35 = El Yunque, near 900 m elevation on transect trail (DICE), 36 = Phase 2 site along the uphill side of gated forest service road from 186 to the old Espiritu Santo bridge. (CYBO, CYHO, ASSE)

ACRDAN = Acrostichum danaeaifolium, BLEOCC = Blechnum occidentale, BOLNIC = Bolbitis nicotianafolia, CYABOR = Cyathea borinquena, DANEL1 = Danaea elliptica (form 1, Sonadora watershed), DANEL2 = Danaea elliptica (form 2, El Spiritu Santo watershed), DIPCEN = Diplazium centrepetale, DANNOD = Danaea nodosa, ELACRI = Elaphoglossum crinitum, ELASIM = Elaphoglossum simplex (found in Sonadora watershed), ELALAT = Elaphoglossum latifolium (formerly E2), HEMMAR = Hemidictyum marginatum, LINLAN = Lindsaea lancea, LONHIR = Lonchitis hirsuta, NEPRIV = Nephrolepis rivularis, POLCER = Polybotrya cervina (= OLFCER Olfersia cervina), THEANG = Thelypteris angustifolia, THEDEL = Thelypteris deltoidea, THERET = Thelypteris reticulata, TECTRI = Tectaria trifoliata, ASPSER = Asplenium serratum, DIPUNI = Diplazium unilobum, ASPCUN = Asplenium cuneatum, POLCRA = Polypodium crassifolium, ELARIG = Elaphoglossum rigidum, CYAHOR = Cyathea horrida

DATA TYPE

alphanumeric

integer

integer

alphanumeric

MISSING DATA CODES

Must be present

Must be present

Must be present

Must be present

10 File Name or # above (all in which the variable appears)

1-22

1-15, 20

1-15, 20

15, 19, 21, 22

Abbreviation (as it appears on the data file)

SPCODE

STAT

SN

SI

NAME OF VARIABLE

Species code, abbreviated

Status of leaf

Status of leaf modifier

Sub-ID of plant

DEFINITION OF VARIABLE

Code for scientific name of the species of fern, abbreviated to two letters GS (usually genus and species initial) to save space in files

Maturation status of leaf

Additional information about the maturation status of the leaf.

A letter designation indicating either that 1) the plant is too young for a tag to be attached (A, B, C, D, etc.) or is a branch of the parent number, or 2) the plant was considered to be one plant at an earlier time, but is now considered to be two or more plants (S) or 3) the continuity of monitoring was broken for some reason and the plant was re-profiled (S) (Such records are archived, and the sub-id not used in continuation of monitoring).Though the column remains, note that all Sub-ID codes were eliminated and new numbers starting with 9000 were assigned in 2006 as noted in the FERNFLD files to facilitate data analysis.

Note SI still appears on FERNTHAN because all workfiles for that publication include SI.

UNIT

PRECISION

RANGE OR LIST OF VALUES

AD = Acrostichum danaeaifolium, BO = Blechnum occidentale, BN = Bolbitis nicotianafolia, CB = Cyathea borinquena, DE1 = Danaea elliptica (form 1, Sonadora watershed - exception to GS), D2 = Danaea elliptica (form 2, El Spiritu Santo watershed - exception to GS), DC = Diplazium centrepetale, DN = Danaea nodosa, EC = Elaphoglossum crinitum, ES = Elaphoglossum simplex (found in Sonadora watershed), EL = Elaphoglossum latifolium (formerly E2), HM = Hemidictyum marginatum, LL = Lindsaea lancea, LH = Lonchitis hirsuta, NR = Nephrolepis rivularis, PC = Polybotrya cervina (= OLFCER Olfersia cervina), TA = Thelypteris angustifolia, TD = Thelypteris deltoidea, TR = Thelypteris reticulata, TT = Tectaria trifoliata, AS = Asplenium serratum, DU = Diplazium unilobum, AC = Asplenium cuneatum, PC = Polypodium crassifolium, ER = Elaphoglossum rigidum, CH = Cyathea horrida

T = Fully expanded leaf (was M in earlier field notes) mature leaf,
I = Immature; not fully expanded (if no change in leaf length in subsequent monitoring period, may be changed to T. If both I and B, then enter I and put B in the gone code.), B = Broken leaf (or cut if C follows In SN field. If both I (immature) and B, then enter I and put B in the gone code.), C = crozier

Broken leaf (status B): n =number of pairs left, 0 = broken tip, blank or 99 = only petiole left

Crozier (status C): n = pair of leaflets at top of coil, counting from first (lowermost) pair unrolled, 0 = only tip still coiled

Other codes: LS = sori light in color, H = hairy (ElCr), C = cut by machete (AcDa only)

A - J, S

DATA TYPE

alphanumeric

alphanumeric

alphanumeric

alphanumeric

MISSING DATA CODES

Must be present

Must be present

Blank

11 File Name or # above (all in which the variable appears)

1-15, 20

1-15, 20

1-15, 20

1-15, 20

Abbreviation (as it appears on the data file)

TAG

TYPELEAF

TN

WID

NAME OF VARIABLE

Tagged leaf code

Type of leaf

Type of leaf modifier

Width

DEFINITION OF VARIABLE

A code to indicate that leaf has now or has had in the past a label affixed to it.

Type of leaf in terms of reproductive status

Additional information about the Type code

Width of the leaf at its widest point for NEPRIV and ELACRI prior to 2003. From 2003 on, should be measured for all leaves.

UNIT

centimeter

PRECISION

RANGE OR LIST OF VALUES

L =label attached at some point while leaf monitored, Blank = no label ever attached.

S = Sterile (photosynthetic only; no spores), F = Fertile (sporophyll, with sori present), U = Unknown, C = Crozier, therefore unknown, L = Leaf base only

P = partially fertile (not all leaflets expected to have sori do), T = Sori smaller than usual (THDE), ! = exceptionally robust sori (THDE)
n (1-99) = number of pairs of fertile leaflets (NERI), n (1, 2, 3) = Juvenile, twice-cut or thrice-cut (CYBO)

DATA TYPE

alphanumeric

alphanumeric

alphanumeric or integer

numeric

MISSING DATA CODES

Must be present. Note that for Profile code S or D (croziers), this code will be entered some time after the mature leaf has been observed.

12 File Name or # above (all in which the variable appears)

1-15, 20

1-17, 20

1-15, 20

14

14

Abbreviation (as it appears on the data file)

WIRE

PFERT

WSHED

TRT61

TRT161

NAME OF VARIABLE

Wire color

Plant fertility code

Watershed code

Defoliation experiment code for THDE-Round 1

Defoliation experiment code for THDE-Round 2

DEFINITION OF VARIABLE

Color of the 6-10 cm plastic insulated copper wire attached to leaf when first measured. Used only for leaf identification in the field when all individual leaves are measured.

Indicates whether or not fertile (spore bearing) leaves were ever encountered on the plant.

Code indicating which river's watershed the plant was measured in

Code indicating that the leaf was either defoliated or emerged after a defoliation event.

Code indicating that the leaf was either defoliated or emerged after a defoliation event.

UNIT

N/A

N/A

N/A

N/A

N/A

PRECISION

N/A

N/A

N/A

N/A

N/A

RANGE OR LIST OF VALUES

R = Red, Y = Yellow, B = Blue, G = Green, O = Orange, P = Purple, K = Black, N = Brown, A = Gray, W = White, HP = no wire, hole punched in lamina instead, blank = leaf too immature to attach wire without damage, between colors = two wires; first color replaced second

F = fertile leaves present in crown at some point, S = no fertile leaves ever present in the crown (plant is sterile), Phase2 (file 16) includes only those observations made in the phase 2 format, blank = Can be blank until final monitoring of the plant has been completed

1 = Rio Sonadora (El Verde), 2 = Rio Espiritu Santo, 3 = Rio Mamayes (Bisley), blank = Can be blank until needed for analysis.

TRT61: 1 = defoliated in September 1996, 2 = defoliated in September 1997

TRT161: 1 = defoliated in January 2005

DATA TYPE

alphanumeric

alphabetic

numeric

numeric

numeric

MISSING DATA CODES

N/A

N/A

N/A

Not part of the THDE defoliation experiment

Not part of the THDE defoliation experiment

13 File Name or # above (all in which the variable appears)

14

Abbreviation (as it appears on the data file)

BLEAF

NAME OF VARIABLE

Biomass of leaf

DEFINITION OF VARIABLE

Dry weight of leaf removed from THDE plant for defoliation experiment TRT61.

UNIT

grams

PRECISION

N/A

RANGE OR LIST OF VALUES

DATA TYPE

numeric

MISSING DATA CODES

Leaf not weighed

14 File Name or # above (all in which the variable appears)

16, 17

16

16

16

Abbreviation (as it appears on the data file)

CAT2

MFERT

FBRK

FCTN

NAME OF VARIABLE

Phase 2 plant category

Monitoring date leaf fertility status

Fertile leaves broken code

Fertile leaf count--new leaves

DEFINITION OF VARIABLE

Identifies the circumstances of a plants transition from Phase I to Phase II

Indicates whether or not new fertile leaves were produced during the season or year prior to the monitoring date.

If present, this code indicates that all new fertile leaves are broken

Count of the fertile leaves added since the plant last monitored

UNIT

N/A

N/A

N/A

N/A

PRECISION

N/A

N/A

N/A

N/A

RANGE OR LIST OF VALUES

1 = New plants of new species in Phase II, 2 = New plants of Phase I species, measured using phase II forms (no profile needed), 3 = Phase I plants with low turnover rates (continue to record leaf data as before, though fertile leaves may be missed if a dimorphic species), 4 = Phase I plants with high turnover rates though still identifiable (using phase II forms), 5 = New plants with high turnover rates (Phase II form), 6 = Phase I plants which have branched and/or formed clumps (phase II form), 7 = Phase II plants with high turnover rates that can't be identified from one year to next (are assigned special temporary ID numbers 3000 to 8999), 8 = Annual monitoring begun in January 1999 (THERET)

F = New fertile leaves noted, S = No new fertile leaves noted

B= all fertile leaves on the plant are broken, therefore some leaf measurements could not be made, C = Fertile leaf cut, D = Fertile leaf dead but measurable, H = Fertile leaf too insect damaged to measure, Blank = measurements of the longest fertile leaf should be present unless FCTN is equal to 0

DATA TYPE

numeric

alphabetic

alphabetic

integer

MISSING DATA CODES

Must be present

Must be present

N/A

Must be present unless Plant Gone code is present

15 File Name or # above (all in which the variable appears)

16

16

16

16

Abbreviation (as it appears on the data file)

FCTO

FLAM

FLEN

FPET

NAME OF VARIABLE

Fertile leaf count - old leaves

Fertile lamina length

Fertile leaf length

Fertile leaf petiole length

DEFINITION OF VARIABLE

Count of the fertile leaves already tagged

Measurement of the leaf lamina on the longest unbroken new fertile leaf

Length of the longest unbroken new fertile leaf (*Calculated: FPET+FLAM)

Length of the petiole on the longest unbroken new fertile leaf

UNIT

N/A

centimeter

centimeter

centimeter

PRECISION

N/A

N/A

N/A

N/A

RANGE OR LIST OF VALUES

N/A

N/A

N/A

N/A

DATA TYPE

integer

integer

integer

integer

MISSING DATA CODES

Must be present

N/A

N/A

N/A

16 File Name or # above (all in which the variable appears)

16

16

17

16

Abbreviation (as it appears on the data file)

FPRS

FWID

MET

PF

NAME OF VARIABLE

Fertile leaflet pair count

Fertile leaf width

Metal tag number

Plant first monitored

DEFINITION OF VARIABLE

Count of the pairs of leaflets (if present) on the longest unbroken new fertile leaf

Width of the leaf lamina measured at about midpoint of the longest unbroken new fertile leaf

Number of the metal "race-track" shaped tag that has been looped around the base of the plant with grafting tape, if present

Very first time plant is measured if plant is first measured in Phase II. All leaf measurements will be "old" as age of plant can't be determined.

UNIT

N/A

centimeter

N/A

N/A

PRECISION

N/A

N/A

N/A

N/A

RANGE OR LIST OF VALUES

N/A

N/A

N/A

P = first time measured, Blank = not first time measured

DATA TYPE

integer

integer

integer

alphabetic

MISSING DATA CODES

N/A

N/A

N/A

N/A

17 File Name or # above (all in which the variable appears)

16, 17

16

16

Abbreviation (as it appears on the data file)

PGCODE

SBRK

SCTN

NAME OF VARIABLE

Plant gone code

Sterile leaves broken code

Sterile leaf count--new leaves

DEFINITION OF VARIABLE

Code used to indicate why the plant is no longer being monitored

If present, this code indicates that all new sterile leaves are broken

Count of the sterile leaves added since the plant last monitored

UNIT

N/A

N/A

N/A

PRECISION

N/A

N/A

N/A

RANGE OR LIST OF VALUES

G = plant gone from a known location and assumed dead, D = plant dead, E = last monitoring of plant was done in January 2001 (end of Phase I) because site was dropped (5, 10, Jobos Bay). May or may not have been dead at that time, so can't be used to determine date of death, X = monitoring stopped for reasons described in field notes; can not assume plant death. For example, for a clumped plant, can't tell individuals after a year, may be alive and monitored under another ID as noted in field notes, so not a "dead" plant code. Or, leaf turnover so rapid that ID tag and/or wires can not be used to identify the plant, etc.

Phase 2 file only: R = plant transferred back to Phase 1 protocol, N = plant not found this year but relocated later, T = temporary ID for one year only (THEANG)

B = all sterile leaves on plant are broken, therefore no leaf measurements could be made, C = cut sterile leaves so no leaf measurement could be made, Blank = measurements of the longest sterile leaf should be present unless SCTN is equal to 0

N/A

DATA TYPE

alphabetic

alphabetic

integer

MISSING DATA CODES

N/A

N/A

Must be present

18 File Name or # above (all in which the variable appears)

16

16

16

16

Abbreviation (as it appears on the data file)

SCTO

SLAM

SLEN

SPET

NAME OF VARIABLE

Sterile leaf count - old leaves

Sterile leaf lamina length

Sterile leaf length

Sterile leaf petiole length

DEFINITION OF VARIABLE

Count of sterile leaves already tagged

Measurement of the leaf lamina on the longest unbroken new sterile leaf

Length of the longest unbroken new sterile leaf (*Calculated: SPET+SLAM)

Length of the petiole on the longest unbroken new sterile leaf

UNIT

N/A

centimeter

centimeter

centimeter

PRECISION

N/A

N/A

N/A

N/A

RANGE OR LIST OF VALUES

N/A

N/A

N/A

N/A

DATA TYPE

integer

integer

integer

integer

MISSING DATA CODES

Must be present

N/A

N/A

N/A

19 File Name or # above (all in which the variable appears)

16

16

16

17

Abbreviation (as it appears on the data file)

SPRS

SWID

TOT

PTOT

NAME OF VARIABLE

Sterile leaflet pair count

Fertile leaf width

Total number of leaves in crown

Total leaf count when the plant is first measured

DEFINITION OF VARIABLE

Count of the pairs of leaflets (if present) on the longest unbroken new sterile leaf

Width of the leaf lamina measured at about midpoint of the longest unbroken new sterile leaf

Number of mature leaves, both new and old, in the crown (Phase 2) (*Calculated: FCTO+FCTN+SCTO+SCTN)

Count of leaves measured when plant is profiled, taken from files 1-15, 20 for Phase 1 plants only

UNIT

N/A

centimeter

N/A

N/A

PRECISION

N/A

N/A

N/A

N/A

RANGE OR LIST OF VALUES

N/A

N/A

N/A

N/A

DATA TYPE

integer

integer

integer

integer

MISSING DATA CODES

N/A

N/A

N/A

N/A

20 File Name or # above (all in which the variable appears)

17

17

17

17

Abbreviation (as it appears on the data file)

PFCT

PIM

EFCONSCM

EFSPCODE

NAME OF VARIABLE

Fertile leaf count when plant first measured

Immature leaf count when plant first measured

Conspecific fern distance

Species of nearest fern

DEFINITION OF VARIABLE

Count of fertile leaves in crown when plant is profiled, taken from files 1-15, 20 for Phase 1 plants only

Count of immature leaves and croziers in crown when plant is profiled, taken from files 1-15, 20 for Phase 1 plants

Distance from plant being studied to nearest plant of the same species.

Abbreviated code for species (see SPCODE) of fern nearest plant being studied that is not the same species

UNIT

N/A

N/A

centimeter

N/A

PRECISION

N/A

N/A

N/A

N/A

RANGE OR LIST OF VALUES

N/A

N/A

N/A

AD = Acrostichum danaeaifolium, BO = Blechnum occidentale, BN = Bolbitis nicotianafolia, CB = Cyathea borinquena, D1 = Danaea elliptica (form 1, Sonadora watershed), D2 = Danaea elliptica (form 2, El Spiritu Santo watershed), DC = Diplazium centrepetale, DN = Danaea nodosa, EC = Elaphoglossum crinitum, ES = Elaphoglossum simplex (found in Sonadora watershed), EL = Elaphoglossum latifolium (formerly E2), HM = Hemidictyum marginatum, LL = Lindsaea lancea, LH = Lonchitis hirsuta, NR = Nephrolepis rivularis, PC = Polybotrya cervina (= OLFCER Olfersia cervina), TA = Thelypteris angustifolia, TD = Thelypteris deltoidea, TR = Thelypteris reticulata, TT = Tectaria trifoliata, AS = Asplenium serratum, DU = Diplazium unilobum, AC = Asplenium cuneatum, PC = Polypodium crassifolium, ER = Elaphoglossum rigidum, CH = Cyathea horrida

DATA TYPE

integer

integer

integer

alphabetic

MISSING DATA CODES

N/A

N/A

N/A

Missing for plants gone from study before May 1997

21 File Name or # above (all in which the variable appears)

17

17

17

17

Abbreviation (as it appears on the data file)

EPDM

ESLOPE

ESUB

LOC1, 2, 3

NAME OF VARIABLE

Palm distance

Slope of substrate

Substrate

Location of plant

DEFINITION OF VARIABLE

Distance from plant being studied to nearest Sierra Palm individual (Prestoea montana)

Substrate slope measured as an angle where 0 = horizontal and 90 is vertical

Type of substrate plant is located on

Specific location information for a plant, if needed. May be distance in meter from a known location or general area of a site.

UNIT

meter

degree

N/A

N/A

PRECISION

N/A

N/A

N/A

N/A

RANGE OR LIST OF VALUES

N/A

0-90

PR = Palm root, PS = Palm stump, PT = Palm trunk, R = Rock, RC = Rock crevice, S= Soil, TR = Tree root, TT = Tree trunk

N/A

DATA TYPE

integer

integer

alphabetic

N/A

MISSING DATA CODES

Missing from plants gone from study before May 1997

Missing from plants gone from study before 1997

Missing from plants gone from study before 1997

If missing, not needed for plant.

22 File Name or # above (all in which the variable appears)

17

17

17

17

Abbreviation (as it appears on the data file)

LyymP

LyymT

LyymAL

PSEQ

NAME OF VARIABLE

Litter percent

Litter type

Littered plant part

Plant sequence

DEFINITION OF VARIABLE

Percent of plant covered with litter during the monitoring period noted in column heading (eg: yym = 985 = May 1998)

Type of litter that is on the plant at the time of monitoring

Section of plant that had litter on it. Used only from May 1993 through September 1994 as it became clear that almost all litter fell on the apex.

The sequence in which plants were most easily monitored in the field, irrespective of plant ID number.

UNIT

Percent

N/A

N/A

N/A

PRECISION

N/A

N/A

N/A

N/A

RANGE OR LIST OF VALUES

1-100

CL = Cecropia leaves, CP = Cecropia leaf petiole, L = Leaf litter other than palm, heliconia or cecropia, N = No litter, PR = Palm rachis, PI = Palm inflorescence, PP = Palm petiole, PL = Palm leaflets, R or RT = Root, RK = Rock, S = Stick (small diameter), V = Vine, W = Wood (larger diameter)

A = Apex, L = Leaves

N/A

DATA TYPE

integer

alphabetic

alphabetic

numeric

MISSING DATA CODES

N/A

N/A

N/A

Must be present if plant is being actively monitored

23 File Name or # above (all in which the variable appears)

17

Abbreviation (as it appears on the data file)

ORIGIN

NAME OF VARIABLE

Plant origin

DEFINITION OF VARIABLE

Plant origin if known to have branch off of another plant in the study.

UNIT

N/A

PRECISION

RANGE OR LIST OF VALUES

X = branched from another plant

DATA TYPE

alphanumeric

MISSING DATA CODES

N/A

24 File Name or # above (all in which the variable appears)

18, 19, 21, 22

19, 21, 22

19, 21, 22

18, 19, 21 22

Abbreviation (as it appears on the data file)

FNSOURCE

FNCODE

FNUSE

FPG

NAME OF VARIABLE

Field note source

Field note code

Field note active

Field book page

DEFINITION OF VARIABLE

Field book number, starting at the beginning of the study. May not be present if general notes not recorded in field book.

DS = Notes from data sheets recorded in the field

AN = Notes added during data analysis

Indicates the type of information being noted using an alphabetic code followed bya sequence number if the notes use multiple lines

Any field notes which much be printed on the data sheets for monitoring (ex: plant location information) are noted.

All field books have numbered pages for relocating notes and data entered, blank = notes not recorded in field books

UNIT

N/A

N/A

N/A

N/A

PRECISION

N/A

N/A

N/A

N/A

RANGE OR LIST OF VALUES

1-53 (Phase 1) 54...(Phase 2), DS, AN

Note:Additional codes can be found at the beginning of the FERNFLD files.

A = information on rhizome division and re-identification, B = branch from a main trunk (MT), C = ID changed to remove SubID, E = Leaf information, F = litterFall information, FA = Field Assistant, G = no longer monitored, with reasons given (ex: dead, not found, etc.), H = Herbivory observation, I = Index for page number for site monitored, L = Location, M = mapping data, MT = main trunk when plant rhizome divides, N = general Notes, P = Photography noted, R = Rhizome observations, S = Site notes, T = Trees in area, V = Voucher information, W = Water level information, Z = Zoological observations

F = Field note active, to be used for monitoring data sheets

N/A

DATA TYPE

alphanumeric

alphabetic

alphabetic

numeric

MISSING DATA CODES

N/A

Must be present

N/A

N/A

25 File Name or # above (all in which the variable appears)

19, 21, 22

18, 19, 21, 22

Abbreviation (as it appears on the data file)

FNOTE

YYYYM

NAME OF VARIABLE

Field notes

Year and month

DEFINITION OF VARIABLE

Any type of information noted, generally recordedin the field, that might be useful in locating the plant or interpreting the results. In addition data about plant mortality and various aspects of the leaves and rhizome are noted.

Year and month for the monitoring dates during which the notes were made (note that all monitoring was done in single-digit months).

UNIT

N/A

N/A

PRECISION

N/A

N/A

RANGE OR LIST OF VALUES

N/A

N/A

DATA TYPE

alphanumeric

numeric

MISSING DATA CODES

Must be present

Must be present

26 File Name or # above (all in which the variable appears)

18

18

18

18

Abbreviation (as it appears on the data file)

DATE

FA

FTEND

FTSPAN

NAME OF VARIABLE

Date of monitoring

Field assistant

Field time end

Field time span

DEFINITION OF VARIABLE

Date the site was monitored (MM/DD/YYYY)

Initials of the person who assisted with monitoring, if not alone.

Time the work at the site was begun, if noted.

Total time spent at the site, if noted.

UNIT

N/A

N/A

N/A

Minute

PRECISION

N/A

N/A

N/A

N/A

RANGE OR LIST OF VALUES

CR = Conchi Rodrigues, JH = Jessie Hopkins, JR = Julie Rea, KS = Kathryn Lee Stack Schwertfeger, LF = Leslie Finical Halleck, MW = Megan Weaver, TJB = Thomas Jackson Bennett

N/A

N/A

DATA TYPE

Date

alphabetic

Time

numeric

MISSING DATA CODES

Must be present

N/A

N/A

N/A

27 File Name or # above (all in which the variable appears)

18

18

18

18

Abbreviation (as it appears on the data file)

FSTART

JDATE

MO

SERIALDATE

NAME OF VARIABLE

Field time start

Julian date

Monitoring month

Sequential number assigned to dates starting at January 1, 1900.

DEFINITION OF VARIABLE

Time the work at the site was begun, if noted.

Julian date at site if noted.

Month site was monitored.

Date is converted, using MS Excel or Lotus 1-2-3, to sequential number by changing format from Date to General

UNIT

N/A

N/A

N/A

N/A

PRECISION

N/A

N/A

N/A

N/A

RANGE OR LIST OF VALUES

N/A

1-366

1-12

33491-present

DATA TYPE

Time

numeric

numeric

General

MISSING DATA CODES

N/A

N/A

Must be present

Must be present

28 File Name or # above (all in which the variable appears)

18

Abbreviation (as it appears on the data file)

YR

NAME OF VARIABLE

Monitoring year

DEFINITION OF VARIABLE

Year that site was monitored (YYYY)

UNIT

N/A

PRECISION

N/A

RANGE OR LIST OF VALUES

1991 - present

DATA TYPE

numeric

MISSING DATA CODES

Must be present

Variable

Page

Page

Variable

ADD

FPET

15

MET

16

SI

10

BLEAF

13

FPG

24

MFERT

14

SITE

9

CAT2

14

FPRS

16

MO

27

SLAM

18

COND

FTEND

26

MPHEN

7

SLEN

18

CUTCODE

2

FTOT

23

MS

6

SN

10

CUTMS

2

FTSPAN

26

MYRPHEN

7

SP

9

CUTPET

2

FTSTART

27

OR

7

SPCODE

10

DATE

26

FWID

16

ORIGIN

23

SPET

18

DAY

3

GC

3

PET

8

SPRS

19

DDIF

2

GPHEN

3

PETI

8

STAT

10

DIF

3

GS

4

PF

16

STOT

23

EFCONSCM

20

GYRPHEN

4

PFCT

20

SWID

19

EFSPCODE

20

ID

4

PFERT

12

TAG

11

EPDM

21

JDATE

27

PGCODE

17

TN

11

ESLOPE

21

LAM

4

PIM

20

TOT

19

ESUB

21

LAM1C

5

PR

9

TRT61,161

12

FA

26

LAM2C

5

PRS

7

TYPE

11

FBK

24

LAM2I

5

PRS1

8

WID

11

FBRK

14

LAMII

5

PRS2

8

WIRE

12

FCTN

14

LEAF

6

PSEQ

22

WSHED

12

FCTO

15

LEN

6

PTOT

19

YR

28

FLAM

15

LN

6

SBRK

17

YYYYM

25

FLEN

15

LOC1,2,3

21

SCTN

17

FNCODE

24

LyymAL

22

SCTO

18

FNOTE

25

LyymP

22

SEQ

9

FNUSE

24

LyymT

22

SERIALDATE

27

VARIABLE

FORMULA

FLEN

FPET+FLAM

SLEN

SPET+SLAM

FTOT

FCTO+FCTN

STOT

SCTO+SCTN

TOT

FCTO+FCTN+SCTO+SCTN

LAM1C

Lamina 1/Inches * 2.54

LAM2C

Lamina 2/Inches * 2.54

LEN

Sum of petiole and lamina length.

PET

For Acrostichum, the petiole length is calculated (PETI * 2.54)

PRS

1. Counted in field, usually on the right side with adaxial surface up. If pairs on both side are counted (occasionally done on smaller leaves) and are unequal in number, then .5 is used to average the number on both sides of rachis. 2. For leaves first encountered as croziers (Profile codes S and D), the number of pairs counted when the leaf is mature is inserted in this field later.