Table 1. Research Themes to Be Addressed in LEF-LTER 3.

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Research Theme 1:  Cumulative Effects of Disturbance – Hurricanes Hugo and Georges and Legacies of Human Land Use (Continuation of LTER 2)
Research Theme 2: The Role of the Biota in Controlling Community and Ecosystem Processes that Affect Nutrient Availability and Organic Matter Processing
Research Theme 3:  The Influence of Climate and Physical Constraints on the Distribution and Abundance of Organisms and Related Ecosystem Processes in the Luquillo Mountains

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15. Model for the Luquillo LTER showing the relationship between disturbance, physical template, primary gradients of water, light, and nutrients (abiotic factors), and the biota.