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| Research Theme 1: | Cumulative Effects of Disturbance – Hurricanes Hugo and Georges and Legacies of Human Land Use (Continuation of LTER 2) |
| Research Theme 2: | The Role of the Biota in Controlling Community and Ecosystem Processes that Affect Nutrient Availability and Organic Matter Processing |
| Research Theme 3: | The Influence of Climate and Physical Constraints on the Distribution and Abundance of Organisms and Related Ecosystem Processes in the Luquillo Mountains |
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